Veronica triloba |
Veronica anagallis-aquatica |
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trilobed speedwell |
blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
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Habit | Annuals. | Annuals or perennials. |
Stems | decumbent, (5–)10–30 cm, ± eglandular-hairy. |
erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
Leaves | blade broadly ovate to suborbiculate, 3(–5)-lobed, central lobe 1/4–1/2 times length of whole leaf, (3–)5–10(–15) × (4–)6–12(–18) mm, ± fleshy, base truncate to subcordate, margins 3(–5)-lobed, apex acute, surfaces sparsely eglandular-hairy. |
petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
Racemes | 1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis sparsely eglandular-hairy; bracts broadly ovate to suborbiculate, 3(–5)-lobed, (3–)5–10(–15) mm. |
6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
Pedicels | patent, deflexed in fruit, (2–)4–8(–10) mm, shorter than subtending bract, length 1–2 times calyx, glabrous abaxially, hairy adaxially. |
suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
Flowers | calyx lobes 4–5(–6) mm, apex acute, puberulent on abaxial surface, rarely glabrous, densely ciliate with 35–60 per side; corolla deep blue with white center, 3–5 mm diam.; stamens dark blue, length unknown; style 0.5–0.9(–1.1) mm. |
calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
Capsules | ± round in cross section, ovoid, 2.5–3(–3.5) × 4–5 mm, apex ± emarginate, glabrous. |
slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
Seeds | (1–)4, dull yellowish brown, ellipsoid, often elongate-cuboidal, deeply urn-shaped (elaiosome inside), 2.2–2.8 × 1.5–2.2 mm, 1–1.4 mm thick, cristate to strongly rugose at margins. |
40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
2n | = 18 (Eurasia). |
= 36. |
Veronica triloba |
Veronica anagallis-aquatica |
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Phenology | Flowering (Feb–)Mar–May(–Jun). | Flowering Mar–Sep(–Nov). |
Habitat | Disturbed sites. | Stream margins, ditches, banks, springs, swamps, wet meadows. |
Elevation | 100–200 m. (300–700 ft.) | 0–4000 m. (0–13100 ft.) |
Distribution |
CA; OH; Europe (Balkans); Asia (Turkey) [Introduced in North America] |
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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Discussion | Veronica triloba is known in the flora area from three recent collections in California and Ohio; it is possibly more widespread than reported here. It is closely related to V. hederifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 316. | FNA vol. 17, p. 313. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | V. hederifolia var. triloba | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera |
Name authority | (Opiz) Opiz: Naturalientausch 11: 467. (1826) | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) |
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