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large speedwell

Copeland's speedwell

Habit Perennials. Perennials.
Stems

erect, rarely ascending, (15–)30–70(–100) cm, eglandular-hairy, hairs evenly distributed around stem.

ascending, unbranched, 5–15 cm, densely glandular-hairy.

Leaves

blade ovate or oblong-ovate, rarely lanceolate, (20–)30–55(–70) × (6–)15–25(–45) mm, 2–3(–4) times as long as wide, base ± cordate, margins dentate to serrate to crenate-incised, rarely subentire, apex acute, abaxial surface hairy, adaxial glabrous.

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

Racemes

2–4(–8), axillary, 60–150 mm, 50–150-flowered, axis hairy;

bracts linear-lanceolate to linear, 4–8(–14) mm.

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

Pedicels

erect, 3–7(–14) mm, equal to subtending bract, eglandular-hairy.

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

Flowers

calyx 5-lobed, abaxial lobes 3–4 mm, adaxial 1–1.5 mm, apex acute, glabrous or hairy;

corolla bright blue, rarely pinkish or white, (9–)12–18 mm diam.;

stamens 4.5–6 mm;

style 5–6 mm.

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

Capsules

± compressed to ± round in cross section, broadly obovoid or obcordiform, (3–)4–5.5 × (2.5–)3.5–5 mm, apex not deeply, narrowly emarginate, glabrous or sparsely hairy.

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

Seeds

(2–)8–18, brown, widely ellipsoid to widely obovoid, flat, 1.2–2.1 × 1–1.5 mm, 0.2–0.4 mm thick, smooth.

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

2n

= 48, 64 (Europe).

= 18.

Veronica teucrium

Veronica copelandii

Phenology Flowering May–Jul. Flowering Jul–Aug.
Habitat Dry meadows, forest edges, scrub, calcareous soils. Subalpine meadows, alpine slopes.
Elevation 0–1000 m. (0–3300 ft.) 1900–2500 m. (6200–8200 ft.)
Distribution
from FNA
CT; DC; IL; IN; MA; MD; ME; MI; MO; NH; NJ; NY; OH; PA; SD; VT; WI; NF; ON; QC; Europe [Introduced in North America]
from FNA
CA
[BONAP county map]
Discussion

Veronica teucrium has often been considered an infraspecific taxon of V. austriaca; many records of V. teucrium have been reported under the name V. austriaca. The occurrence of V. austriaca in the narrow sense in the flora area could be neither supported nor excluded; it differs from V. teucrium mainly in distal leaves of the vegetative shoot linear and entire.

Related species have been introduced for horticultural reasons but apparently have not escaped. Veronica satureiifolia Poiteau & Turpin was reported from Missouri by G. Yatskievych [1999–2013; as V. scheereri (J.-P. Brandt) Holub] outside cultivation based on a single specimen and was similarly found in MIN but likely not naturalized. The name V. latifolia Linnaeus, which has at times been used for V. teucrium, is now considered ambiguous and should not be used (R. K. Brummitt 2007). Reports of V. teucrium in Saskatchewan have not been verified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 322. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. austriaca subsp. teucrium, V. austriaca var. teucrium
Name authority Linnaeus: Sp. Pl. ed. 2, 1: 16. (1762) Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi)
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