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spike speedwell, véronique en épi

undulate speedwell, wavy-leaf water speedwell

Habit Perennials. Annuals or perennials.
Stems

ascending, usually with 4–10 nodes, 5–45(–60) cm, proximally eglandular- or glandular-hairy, sometimes glabrous, distally eglandular- and, usually, glandular-hairy.

erect or prostrate basally, 10–100 cm, glabrous at least proximally, glandular-hairy distally.

Leaves

blade ovate-oblong to narrowly oblong-lanceolate, (30–)40–70(–80) × 5–20 mm, base long-cuneate, margins shallowly crenate to shallowly serrate to subentire, apex obtuse to rounded, surfaces densely glandular- or eglandular-hairy, rarely glabrate.

petiole 0–5 mm;

blade elliptic to ovate, sometimes ovate-oblong or linear-lanceolate, rarely lanceolate, 20–50(–100) × 5–20(–25) mm, 2.5–4 times as long as wide, base attenuate, upwards amplexicaul, margins subentire or crenate to serrate or ± undulate, apex acute to obtuse, surfaces glabrous.

Racemes

1(–7), terminal, sometimes with lateral ones, (50–)70–130(–170) mm, (50–)100–300-flowered, axis eglandular- and glandular-hairy;

bracts oblong to lanceolate, 3–5 mm.

6–25, axillary, 50–220 mm, 10–15 mm diam., 10–100-flowered, axis sparsely glandular-hairy, rarely glabrous;

pedicels 2–6 per cm;

bracts linear to lanceolate, 3–6 mm, apex acute.

Pedicels

suberect to patent, 0.5–1(–2) mm, shorter than subtending bract, eglandular- and glandular-hairy.

patent, 3–5 mm, equal to subtending bract, sparsely glandular-hairy.

Flowers

calyx lobes 2–3 mm, ciliate, apex obtuse, glandular-hairy, rarely eglandular-hairy;

corolla blue, campanulate, longer than wide, 5–6 mm diam., lobes 3–4.5 × 1.5–2 mm;

stamens 5 mm;

style 8 mm.

calyx lobes 1.5–2(–3) mm, apex acute, sparsely glandular-hairy, rarely glabrous;

corolla pale blue or pale lilac, rarely white, 2.5–5 mm diam.;

stamens 2 mm;

style 0.9–2 mm.

Capsules

± compressed in cross section, broadly ovoid, 2–3 × 2–3 mm, apex obtuse, densely glandular-puberulent.

slightly compressed in cross section, obcordiform to globular, 2–3 × 2–3 mm, apex rounded or ± emarginate, sparsely glandular-hairy.

Seeds

3–40, light brown, ellipsoid, flat, 0.6–1.2 × 0.5–0.8 mm, 0.2–0.4 mm thick, smooth.

30–40, ochre, broadly ellipsoid, ± flat, convex on both sides, 0.5–0.6 × 0.4–0.5 mm, thickness and texture unknown.

2n

= 34, 68 (Europe).

= 54 (Asia).

Veronica spicata

Veronica undulata

Phenology Flowering Jun–Sep. Flowering (Mar–)Apr–Sep.
Habitat Dry grasslands. Disturbed, wet places, ditches, or swamps.
Elevation 0–700 m. (0–2300 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
CT; NH; NY; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
AL; OR; WA; Asia [Introduced in North America]
[BONAP county map]
Discussion

Veronica spicata is widely distributed in horticulture and a multitude of cultivars is available. Some specimens may also be derived from a related species, V. barrelieri Schott ex Roemer & Schultes, differentiated from V. spicata by glabrous calyx lobes, and in its typical variety, by eglandular pubescence. Another closely related species, V. incana Linnaeus, has a dense white woolly indumentum and is native to northern Asia and eastern Europe; it has not escaped from cultivation in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the flora area, Veronica undulata was introduced near ports (Mobile, Alabama, and Portland, Oregon) before 1900 via ship ballast from trade with Asia, did not spread much, and may not have persisted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 308. FNA vol. 17, p. 314.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. verna, V. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 10. (1753) Wallich: in W. Roxburgh, Fl. Ind. 1: 147. (1820)
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