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spike speedwell, véronique en épi

ivy speedwell, ivy-leaf speedwell

Habit Perennials. Annuals.
Stems

ascending, usually with 4–10 nodes, 5–45(–60) cm, proximally eglandular- or glandular-hairy, sometimes glabrous, distally eglandular- and, usually, glandular-hairy.

decumbent to ascending, 5–40(–50) cm, eglandular-hairy.

Leaves

blade ovate-oblong to narrowly oblong-lanceolate, (30–)40–70(–80) × 5–20 mm, base long-cuneate, margins shallowly crenate to shallowly serrate to subentire, apex obtuse to rounded, surfaces densely glandular- or eglandular-hairy, rarely glabrate.

blade suborbiculate, (3–)5-lobed, central lobe usually overtopping lateral ones, (5–)7–15(–20) × 8–16(–25) mm, base truncate, margins (3–)5-lobed, apex acute, surfaces sparsely eglandular-hairy.

Racemes

1(–7), terminal, sometimes with lateral ones, (50–)70–130(–170) mm, (50–)100–300-flowered, axis eglandular- and glandular-hairy;

bracts oblong to lanceolate, 3–5 mm.

1–10, terminal, 50–400(–500) mm, 5–20-flowered, axis eglandular-hairy;

bracts suborbiculate, (3–)5-lobed, central lobe usually overtopping lateral ones, (5–)7–15(–20) mm.

Pedicels

suberect to patent, 0.5–1(–2) mm, shorter than subtending bract, eglandular- and glandular-hairy.

patent or deflexed, (5–)9–15(–20) mm, equal to or ± shorter than subtending bract, length 2–3 times calyx, eglandular-hairy (in single line adaxially).

Flowers

calyx lobes 2–3 mm, ciliate, apex obtuse, glandular-hairy, rarely eglandular-hairy;

corolla blue, campanulate, longer than wide, 5–6 mm diam., lobes 3–4.5 × 1.5–2 mm;

stamens 5 mm;

style 8 mm.

calyx lobes 5–6(–7) mm, apex acute, abaxial surface usually glabrous, ciliate with 25–35 hairs per side;

corolla bright blue with bright white center, 5–7(–9) mm diam.;

stamens sky blue, 0.7–1.2 mm;

style (0.6–)0.7–0.9(–1.1) mm.

Capsules

± compressed in cross section, broadly ovoid, 2–3 × 2–3 mm, apex obtuse, densely glandular-puberulent.

± round in cross section, ovoid, 3–4 × 4.5–6 mm, apex ± emarginate, glabrate.

Seeds

3–40, light brown, ellipsoid, flat, 0.6–1.2 × 0.5–0.8 mm, 0.2–0.4 mm thick, smooth.

1–4, bright yellow, ellipsoid to subglobular, urn-shaped, 2.3–3.3 × 2–3 mm, 1–2.2 mm thick, weakly cristate or rugose, ± smooth.

2n

= 34, 68 (Europe).

= 54 (Eurasia).

Veronica spicata

Veronica hederifolia

Phenology Flowering Jun–Sep. Flowering (Feb–)Mar–Jun.
Habitat Dry grasslands. Fields, lawns, gardens, ruderal places, vineyards, open forests, shady rocky places, dunes.
Elevation 0–700 m. (0–2300 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
CT; NH; NY; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MI; MO; NC; NE; NJ; NY; OH; OK; OR; PA; SC; SD; TN; UT; VA; WA; WV; BC; ON; Eurasia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica spicata is widely distributed in horticulture and a multitude of cultivars is available. Some specimens may also be derived from a related species, V. barrelieri Schott ex Roemer & Schultes, differentiated from V. spicata by glabrous calyx lobes, and in its typical variety, by eglandular pubescence. Another closely related species, V. incana Linnaeus, has a dense white woolly indumentum and is native to northern Asia and eastern Europe; it has not escaped from cultivation in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 308. FNA vol. 17, p. 315.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 10. (1753) Linnaeus: Sp. Pl. 1: 13. (1753) — (as hederaefolia)
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