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spike speedwell, véronique en épi

Cusick's speedwell

Habit Perennials. Perennials.
Stems

ascending, usually with 4–10 nodes, 5–45(–60) cm, proximally eglandular- or glandular-hairy, sometimes glabrous, distally eglandular- and, usually, glandular-hairy.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade ovate-oblong to narrowly oblong-lanceolate, (30–)40–70(–80) × 5–20 mm, base long-cuneate, margins shallowly crenate to shallowly serrate to subentire, apex obtuse to rounded, surfaces densely glandular- or eglandular-hairy, rarely glabrate.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1(–7), terminal, sometimes with lateral ones, (50–)70–130(–170) mm, (50–)100–300-flowered, axis eglandular- and glandular-hairy;

bracts oblong to lanceolate, 3–5 mm.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

suberect to patent, 0.5–1(–2) mm, shorter than subtending bract, eglandular- and glandular-hairy.

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes 2–3 mm, ciliate, apex obtuse, glandular-hairy, rarely eglandular-hairy;

corolla blue, campanulate, longer than wide, 5–6 mm diam., lobes 3–4.5 × 1.5–2 mm;

stamens 5 mm;

style 8 mm.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

± compressed in cross section, broadly ovoid, 2–3 × 2–3 mm, apex obtuse, densely glandular-puberulent.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

3–40, light brown, ellipsoid, flat, 0.6–1.2 × 0.5–0.8 mm, 0.2–0.4 mm thick, smooth.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 34, 68 (Europe).

= 18, 72.

Veronica spicata

Veronica cusickii

Phenology Flowering Jun–Sep. Flowering Jun–Aug.
Habitat Dry grasslands. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 0–700 m. (0–2300 ft.) 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
CT; NH; NY; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica spicata is widely distributed in horticulture and a multitude of cultivars is available. Some specimens may also be derived from a related species, V. barrelieri Schott ex Roemer & Schultes, differentiated from V. spicata by glabrous calyx lobes, and in its typical variety, by eglandular pubescence. Another closely related species, V. incana Linnaeus, has a dense white woolly indumentum and is native to northern Asia and eastern Europe; it has not escaped from cultivation in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 308. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority Linnaeus: Sp. Pl. 1: 10. (1753) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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