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spike speedwell, véronique en épi

bird's-eye speedwell, Germander speedwell, véronique petit-chêne

Habit Perennials. Perennials.
Stems

ascending, usually with 4–10 nodes, 5–45(–60) cm, proximally eglandular- or glandular-hairy, sometimes glabrous, distally eglandular- and, usually, glandular-hairy.

ascending, rarely erect, (7–)10–30(–50) cm, densely eglandular-hairy, hairs along stem in 2 prominent lines.

Leaves

blade ovate-oblong to narrowly oblong-lanceolate, (30–)40–70(–80) × 5–20 mm, base long-cuneate, margins shallowly crenate to shallowly serrate to subentire, apex obtuse to rounded, surfaces densely glandular- or eglandular-hairy, rarely glabrate.

blade narrowly ovate to ovate-orbiculate, (10–)12–30(–42) × (6–)10–22(–30) mm, 1–2 times as long as wide, base truncate to ± cordate, margins crenate to deeply incised, apex obtuse, surfaces variably hairy.

Racemes

1(–7), terminal, sometimes with lateral ones, (50–)70–130(–170) mm, (50–)100–300-flowered, axis eglandular- and glandular-hairy;

bracts oblong to lanceolate, 3–5 mm.

1–4, axillary, 40–100(–200) mm, 15–40(–60)-flowered, axis eglandular-hairy, sometimes also glandular-hairy;

bracts linear-elliptic, 3–7 mm.

Pedicels

suberect to patent, 0.5–1(–2) mm, shorter than subtending bract, eglandular- and glandular-hairy.

suberect, (3–)5–8(–10) mm, equal to or shorter than subtending bract, eglandular- and glandular-hairy.

Flowers

calyx lobes 2–3 mm, ciliate, apex obtuse, glandular-hairy, rarely eglandular-hairy;

corolla blue, campanulate, longer than wide, 5–6 mm diam., lobes 3–4.5 × 1.5–2 mm;

stamens 5 mm;

style 8 mm.

calyx 4-lobed, lobes 2–8 mm, apex acute, eglandular- and, sometimes, glandular-hairy;

corolla blue obscure darker nerves and sometimes whitish margin, (6–)10–14(–17) mm diam.;

stamens 4.5–6.5 mm;

style (2.5–)4–5 mm.

Capsules

± compressed in cross section, broadly ovoid, 2–3 × 2–3 mm, apex obtuse, densely glandular-puberulent.

strongly compressed in cross section, obcordiform to obdeltoid, (2–)3.5–4(–5) × (3.5–)4–5(–5.5) mm, apex ± emarginate, eglandular-hairy.

Seeds

3–40, light brown, ellipsoid, flat, 0.6–1.2 × 0.5–0.8 mm, 0.2–0.4 mm thick, smooth.

(2–)12–20(–28), yellow, ellipsoid, flat, 1.1–1.7 × 0.6–1.5 mm, 0.2–0.4 mm thick, smooth to subrugose.

2n

= 34, 68 (Europe).

= 16, 32 (Eurasia).

Veronica spicata

Veronica chamaedrys

Phenology Flowering Jun–Sep. Flowering Apr–Jun(–Oct).
Habitat Dry grasslands. Rich soils, deciduous forests, forest edges, roadsides, chaparral, scrub, meadows, lawns.
Elevation 0–700 m. (0–2300 ft.) 0–2200 m. (0–7200 ft.)
Distribution
from FNA
CT; NH; NY; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
AK; CT; DC; ID; IL; IN; MA; MD; ME; MI; MO; MT; NC; NH; NJ; NY; OH; OR; PA; RI; VA; VT; WA; WI; WV; AB; BC; NB; NF; NS; ON; PE; QC; Eurasia [Introduced in South America (Argentina)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica spicata is widely distributed in horticulture and a multitude of cultivars is available. Some specimens may also be derived from a related species, V. barrelieri Schott ex Roemer & Schultes, differentiated from V. spicata by glabrous calyx lobes, and in its typical variety, by eglandular pubescence. Another closely related species, V. incana Linnaeus, has a dense white woolly indumentum and is native to northern Asia and eastern Europe; it has not escaped from cultivation in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The description provided here for Veronica chamaedrys is solely for the tetraploid cytotype, most probably the exclusive cytotype in the flora area and in central and western Europe. The diploid cytotype is so far only known from eastern and southeastern Europe (K. E. Bardy et al. 2010). A significant change in morphology can occur in shaded habitats, in which especially the petiole can be elongated beyond the range given.

It is unclear whether Veronica chamaedrys is introduced throughout the flora area; it may be native in northeastern areas of North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 308. FNA vol. 17, p. 318.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 10. (1753) Linnaeus: Sp. Pl. 1: 13. (1753)
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