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thyme-leaf speedwell

alpine speedwell, American alpine speedwell, hairy speedwell, spring speedwell, thyme-leaf speedwell, wormskjold's alpine speedwell

Habit Perennials. Perennials.
Stems

creeping to ascending, 5–40 cm, scattered eglandular hairs only, often also with glandular hairs, sometimes glabrate.

erect or ascending, light grayish green distally, unbranched, (3–)8–50 cm, sparsely to densely villous-hirsute, sometimes also glandular-hairy.

Leaves

blade oblong-lanceolate to ovate, 8–25 × 5–13 mm, 1.5–2.5 times as long as wide, base cuneate, margins subentire or serrulate-crenate, apex rounded to short-acuminate, surfaces glabrate.

blade elliptic to lanceolate or oblong-ovate, 8–40 × 5–20 mm, base cuneate, margins entire, dentate, or serrate, apex short-acuminate, surfaces sparsely to densely villous-hirsute or glabrous.

Racemes

1, terminal, 50–100 mm, 10–30(–60)-flowered, axis eglandular- and glandular-hairy;

bracts oblong, 4–7 mm.

1, terminal, 5–40 mm, to 60(–150) mm in fruit, (2–)5–25-flowered, axis densely villous-hirsute and ± glandular-hairy;

bracts linear to lanceolate, 1–8 mm.

Pedicels

erect, 2–5 mm, 4–6 mm in fruit, shorter than subtending bract in flower, eglandular- and, sometimes, glandular-hairy.

erect, 2–10(–15) mm, ca. equal to subtending bract, densely villous-hirsute or glandular-hairy.

Flowers

calyx lobes 2–3 mm, apex acute, ciliate;

corolla white, blue, or pink, with purple or dark blue veins (except on abaxial lobe), rotate, 5–8 mm diam.;

stamens 2.5–3 mm;

style 2–4 mm.

calyx lobes (2.5–)3–5.5(–7) mm, apex obtuse or acuminate, glabrous or densely glandular-hairy;

corolla deep blue-violet or violet-purple to deep blue, rotate, 3–11 mm diam.;

stamens 1–2 mm;

style 0.8–4(–6) mm.

Capsules

compressed in cross section, obcordiform, 2.5–3.5 × 4–5.5 mm, wider than long, apex acutely emarginate, glandular-ciliate, otherwise glabrate.

compressed in cross section, oblong-obovoid, 4–6(–8) × (2.8–)4–5.5 mm, ca. as long as wide, apex emarginate, usually densely glandular-hairy.

Seeds

50–72, brown, ellipsoid, flat, 0.6–1.1 × 0.4–1 mm, 0.2 mm thick, smooth.

10–50, straw colored, ellipsoid, flat, 0.7–1 × 0.4–1 mm, 0.05–1 mm thick, very minutely striate.

2n

= 14.

Veronica serpyllifolia

Veronica wormskjoldii

Phenology Flowering Apr–Aug.
Habitat River banks, wet places, moist meadows, shady forests.
Elevation 0–3300 m. (0–10800 ft.)
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NL; NS; ON; PE; QC; SK; South America; Mexico (Baja California); Eurasia; Australia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; ME; MT; NH; NM; NV; OR; UT; WA; WY; AB; BC; NT; NU; ON; QC; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Alpine plants of Veronica serpyllifolia with bright blue, larger corollas, and smaller, hairier raceme axes have been treated at various ranks under the epithet humifusa. The characters are labile and show intergradations with the typical plants. Veronica serpyllifolia is widespread; it is not clear where it is native.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Veronica wormskjoldii occurs in two cytotypes, a diploid (2n = 18) in western North America and a tetraploid (2n = 36) in eastern North America and Greenland. As demonstrated by D. C. Albach et al. (2006), the tetraploid is likely to be a hybrid of the diploid V. alpina and V. wormskjoldii. In that publication, species rank was used for simplicity, not as a taxonomic conclusion. However, subsequent morphological analyses in conjunction with preparing this treatment revealed that the large variation in the diploid V. wormskjoldii, also supported by the large number of varieties established within that taxon, make it nearly impossible to differentiate the two North American taxa morphologically. In cases where two taxa are morphologically undifferentiable, but karyologically and genetically distinct and geographically clearly separate, the rank of subspecies seems more appropriate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pedicels 2–5 mm; e North America.
subsp. wormskjoldii
1. Pedicels (2–)5–10(–15) mm; w North America.
subsp. nutans
Source FNA vol. 17, p. 312. FNA vol. 17, p. 309.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna
Subordinate taxa
V. wormskjoldii subsp. nutans, V. wormskjoldii subsp. wormskjoldii
Synonyms V. humifusa, V. serpyllifolia var. decipiens, V. serpyllifolia subsp. humifusa, V. serpyllifolia var. humifusa, V. tenella V. alpina subsp. wormskjoldii, V. alpina var. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 1: 101. (1817) — (as wormskjoldi)
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