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thyme-leaf speedwell

Cusick's speedwell

Habit Perennials. Perennials.
Stems

creeping to ascending, 5–40 cm, scattered eglandular hairs only, often also with glandular hairs, sometimes glabrate.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade oblong-lanceolate to ovate, 8–25 × 5–13 mm, 1.5–2.5 times as long as wide, base cuneate, margins subentire or serrulate-crenate, apex rounded to short-acuminate, surfaces glabrate.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1, terminal, 50–100 mm, 10–30(–60)-flowered, axis eglandular- and glandular-hairy;

bracts oblong, 4–7 mm.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

erect, 2–5 mm, 4–6 mm in fruit, shorter than subtending bract in flower, eglandular- and, sometimes, glandular-hairy.

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes 2–3 mm, apex acute, ciliate;

corolla white, blue, or pink, with purple or dark blue veins (except on abaxial lobe), rotate, 5–8 mm diam.;

stamens 2.5–3 mm;

style 2–4 mm.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

compressed in cross section, obcordiform, 2.5–3.5 × 4–5.5 mm, wider than long, apex acutely emarginate, glandular-ciliate, otherwise glabrate.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

50–72, brown, ellipsoid, flat, 0.6–1.1 × 0.4–1 mm, 0.2 mm thick, smooth.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 14.

= 18, 72.

Veronica serpyllifolia

Veronica cusickii

Phenology Flowering Apr–Aug. Flowering Jun–Aug.
Habitat River banks, wet places, moist meadows, shady forests. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 0–3300 m. [0–10800 ft.] 1500–3200 m. [4900–10500 ft.]
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NL; NS; ON; PE; QC; SK; South America; Mexico (Baja California); Eurasia; Australia
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from FNA
CA; ID; MT; OR; WA; BC
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Discussion

Alpine plants of Veronica serpyllifolia with bright blue, larger corollas, and smaller, hairier raceme axes have been treated at various ranks under the epithet humifusa. The characters are labile and show intergradations with the typical plants. Veronica serpyllifolia is widespread; it is not clear where it is native.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 312. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. humifusa, V. serpyllifolia var. decipiens, V. serpyllifolia subsp. humifusa, V. serpyllifolia var. humifusa, V. tenella V. allenii
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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