Veronica serpyllifolia |
Veronica anagallis-aquatica |
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thyme-leaf speedwell |
blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
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Habit | Perennials. | Annuals or perennials. |
Stems | creeping to ascending, 5–40 cm, scattered eglandular hairs only, often also with glandular hairs, sometimes glabrate. |
erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
Leaves | blade oblong-lanceolate to ovate, 8–25 × 5–13 mm, 1.5–2.5 times as long as wide, base cuneate, margins subentire or serrulate-crenate, apex rounded to short-acuminate, surfaces glabrate. |
petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
Racemes | 1, terminal, 50–100 mm, 10–30(–60)-flowered, axis eglandular- and glandular-hairy; bracts oblong, 4–7 mm. |
6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
Pedicels | erect, 2–5 mm, 4–6 mm in fruit, shorter than subtending bract in flower, eglandular- and, sometimes, glandular-hairy. |
suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
Flowers | calyx lobes 2–3 mm, apex acute, ciliate; corolla white, blue, or pink, with purple or dark blue veins (except on abaxial lobe), rotate, 5–8 mm diam.; stamens 2.5–3 mm; style 2–4 mm. |
calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
Capsules | compressed in cross section, obcordiform, 2.5–3.5 × 4–5.5 mm, wider than long, apex acutely emarginate, glandular-ciliate, otherwise glabrate. |
slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
Seeds | 50–72, brown, ellipsoid, flat, 0.6–1.1 × 0.4–1 mm, 0.2 mm thick, smooth. |
40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
2n | = 14. |
= 36. |
Veronica serpyllifolia |
Veronica anagallis-aquatica |
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Phenology | Flowering Apr–Aug. | Flowering Mar–Sep(–Nov). |
Habitat | River banks, wet places, moist meadows, shady forests. | Stream margins, ditches, banks, springs, swamps, wet meadows. |
Elevation | 0–3300 m. [0–10800 ft.] | 0–4000 m. [0–13100 ft.] |
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NL; NS; ON; PE; QC; SK; South America; Mexico (Baja California); Eurasia; Australia
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AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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Discussion | Alpine plants of Veronica serpyllifolia with bright blue, larger corollas, and smaller, hairier raceme axes have been treated at various ranks under the epithet humifusa. The characters are labile and show intergradations with the typical plants. Veronica serpyllifolia is widespread; it is not clear where it is native. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 312. | FNA vol. 17, p. 313. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | V. humifusa, V. serpyllifolia var. decipiens, V. serpyllifolia subsp. humifusa, V. serpyllifolia var. humifusa, V. tenella | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera |
Name authority | Linnaeus: Sp. Pl. 1: 12. (1753) | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) |
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