Veronica peregrina |
Veronica officinalis |
Veronica copelandii |
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neckweed, purslane speedwell, véronique voyageuse |
common gypsyweed, common speedwell, heath speedwell, herbal speedwell, Paul's betony, véronique officinale |
Copeland's speedwell |
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Habit | Annuals. | Perennials. | Perennials. |
Stems | erect or ascending, (2.5–)4–25(–35) cm, glabrous or densely glandular-hairy. |
creeping, decumbent to ascending distally, 10–40(–50) cm, densely eglandular-hairy, rarely glabrate or glabrous. |
ascending, unbranched, 5–15 cm, densely glandular-hairy. |
Leaves | blade oblanceolate proximally, narrowly oblong distally, 5–28(–35) × 2–6(–10) mm, 3–10 times as long as wide, base cuneate, margins entire or dentate distally, apex acute, surfaces glabrous or densely glandular-hairy. |
blade ovate, ovate-oblong, or obovate, (8–)20–35(–50) × (3–)10–20(–30) mm, 1.5–2 times as long as wide, base attenuate to cuneate or obtuse, margins dentate, denticulate, or serrulate, apex obtuse to rounded, surfaces sparsely to densely eglandular-hairy, rarely glabrate. |
blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy. |
Racemes | 1, terminal, sometimes 1 or 2 axillary, 20–200 mm, (2–)5–40-flowered, axis glabrous or densely glandular-hairy; bracts spatulate to linear-lanceolate, 3–22 mm. |
1–4(–8), axillary, 40–60 mm, 10–35(–40)-flowered, axis densely to sparsely eglandular-hairy, sometimes also glandular-hairy, rarely glabrate; bracts linear-lanceolate or narrowly ovate, (1.5–)3–4(–5) mm. |
1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy; bracts lanceolate, 3–5 mm. |
Pedicels | erect, 0.2–1(–2) mm, much shorter than subtending bract, glabrous or densely glandular-hairy. |
erect, (0.5–)1–2(–3) mm, shorter than subtending bract, densely eglandular-hairy, rarely glandular-hairy or glabrate. |
erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy. |
Flowers | calyx lobes (2–)3–6 mm, 0.9–2 mm wide, apex obtuse to acute, glabrous or densely glandular-hairy; corolla white or pale pink, 2–5 mm diam.; stamens 1 mm; style 0.1–0.5 mm. |
calyx 4(or 5)-lobed, lobes 2–3.5(–4) mm, apex obtuse, glandular- and eglandular-hairy; corolla pale blue-lilac to pale pink, rarely white with pink veins, 6–8 mm diam.; stamens 2.8–3.4 mm; style 2.5–3(–4) mm. |
calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy; corolla pale blue to purple, rotate, 8–10 mm diam.; stamens 4–5 mm; style 7 mm. |
Capsules | strongly compressed in cross section, obcordiform, 2.5–5 × 2.5–6 mm, apex ± emarginate, glabrous. |
compressed in cross section, triangulate-obdeltoid to obcordiform, 3–5 × 4–5 mm, apex rounded or truncate, rarely emarginate, glandular-hairy, rarely glabrous. |
compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy. |
Seeds | 12–140, yellow or pale brown, oblong, flat, 0.6–1.6 × 0.4–0.9 mm, 0.1–0.2 mm thick, smooth. |
9–24, brown to yellow, ellipsoid, ovoid, or obovoid, flat, 0.8–1.6 × (0.3–)0.6–1.2 mm, 0.2–0.3 mm thick, smooth. |
number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown. |
2n | = 52. |
= 18, 36 (Europe). |
= 18. |
Veronica peregrina |
Veronica officinalis |
Veronica copelandii |
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Phenology | Flowering (Feb–)Mar–Jun(–Nov). | Flowering (Apr–)Jun–Jul(–Aug). | Flowering Jul–Aug. |
Habitat | Moist waste lands, gardens, roadsides, stream banks, pond shorelines, vernal pools, other cultivated land. | Forests, meadows, pastures, scrub, ruderal places. | Subalpine meadows, alpine slopes. |
Elevation | 0–700(–3000) m. [0–2300(–9800) ft.] | 0–1600 m. [0–5200 ft.] | 1900–2500 m. [6200–8200 ft.] |
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Central America; South America; Mexico (Baja California, Baja California Sur, Chihuahua, México, Sonora); Eurasia; Australia
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CA; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Eurasia [Introduced in North America; introduced in s South America (Chile)]
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CA |
Discussion | Stem indument of Veronica peregrina has been used to distinguish var. xalapensis with a glandular-hairy stem (F. W. Pennell 1935), a variety that seems to be confined to drier places than the type variety. Variety laurentiana is also differentiated from var. peregrina in having a glandular-hairy stem, fleshier leaves, shorter stamens, smaller corollas, and slightly differently shaped capsules (Frère Marie-Victorin and J. Rousseau 1940). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica ×tournefortii Villars [V. officinalis var. tournefortii (Villars) Dumortier] refers to the hybrid between V. officinalis and V. allionii Villars in Europe and is not applicable to any North American plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 314. | FNA vol. 17, p. 311. | FNA vol. 17, p. 310. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | |||
Synonyms | V. peregrina var. laurentiana, V. peregrina subsp. xalapensis, V. peregrina var. xalapensis, V. sherwoodii, V. xalapensis | ||
Name authority | Linnaeus: Sp. Pl. 1: 14. (1753) | Linnaeus: Sp. Pl. 1: 11. (1753) | Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) |
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