Veronica peregrina |
Veronica officinalis |
Veronica americana |
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neckweed, purslane speedwell, véronique voyageuse |
common gypsyweed, common speedwell, heath speedwell, herbal speedwell, Paul's betony, véronique officinale |
American brooklime, American speedwell, véronique d'amérique |
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Habit | Annuals. | Perennials. | Perennials. |
Stems | erect or ascending, (2.5–)4–25(–35) cm, glabrous or densely glandular-hairy. |
creeping, decumbent to ascending distally, 10–40(–50) cm, densely eglandular-hairy, rarely glabrate or glabrous. |
decumbent or ascending, 5–50 cm, glabrous. |
Leaves | blade oblanceolate proximally, narrowly oblong distally, 5–28(–35) × 2–6(–10) mm, 3–10 times as long as wide, base cuneate, margins entire or dentate distally, apex acute, surfaces glabrous or densely glandular-hairy. |
blade ovate, ovate-oblong, or obovate, (8–)20–35(–50) × (3–)10–20(–30) mm, 1.5–2 times as long as wide, base attenuate to cuneate or obtuse, margins dentate, denticulate, or serrulate, apex obtuse to rounded, surfaces sparsely to densely eglandular-hairy, rarely glabrate. |
petiole 2–6(–10) mm; blade ovate to lanceolate, widest proximal to middle, (5–)30–50(–100) × (3–)7–20(–30) mm, 2–4 times as long as wide, base truncate, rounded or almost cordate, abruptly turning into petioles, margins entire or serrate, apex obtuse-orbicular or acute, surfaces glabrous. |
Racemes | 1, terminal, sometimes 1 or 2 axillary, 20–200 mm, (2–)5–40-flowered, axis glabrous or densely glandular-hairy; bracts spatulate to linear-lanceolate, 3–22 mm. |
1–4(–8), axillary, 40–60 mm, 10–35(–40)-flowered, axis densely to sparsely eglandular-hairy, sometimes also glandular-hairy, rarely glabrate; bracts linear-lanceolate or narrowly ovate, (1.5–)3–4(–5) mm. |
1–8, axillary, 30–100 mm, (3–)10–30-flowered, axis glabrous; bracts lanceolate, 3–6 mm. |
Pedicels | erect, 0.2–1(–2) mm, much shorter than subtending bract, glabrous or densely glandular-hairy. |
erect, (0.5–)1–2(–3) mm, shorter than subtending bract, densely eglandular-hairy, rarely glandular-hairy or glabrate. |
patent, 3–10(–12) mm, equal to ± longer than subtending bract, glabrous. |
Flowers | calyx lobes (2–)3–6 mm, 0.9–2 mm wide, apex obtuse to acute, glabrous or densely glandular-hairy; corolla white or pale pink, 2–5 mm diam.; stamens 1 mm; style 0.1–0.5 mm. |
calyx 4(or 5)-lobed, lobes 2–3.5(–4) mm, apex obtuse, glandular- and eglandular-hairy; corolla pale blue-lilac to pale pink, rarely white with pink veins, 6–8 mm diam.; stamens 2.8–3.4 mm; style 2.5–3(–4) mm. |
calyx lobes 2–5(–6) mm, apex acute, glabrous; corolla pink or sky blue, 4–10 mm diam.; stamens 2–4 mm; style (1.7–)2.5–3.5(–4) mm. |
Capsules | strongly compressed in cross section, obcordiform, 2.5–5 × 2.5–6 mm, apex ± emarginate, glabrous. |
compressed in cross section, triangulate-obdeltoid to obcordiform, 3–5 × 4–5 mm, apex rounded or truncate, rarely emarginate, glandular-hairy, rarely glabrous. |
slightly compressed in cross section, globular, 2–4 × 3–5 mm, apex not or ± emarginate, glabrous. |
Seeds | 12–140, yellow or pale brown, oblong, flat, 0.6–1.6 × 0.4–0.9 mm, 0.1–0.2 mm thick, smooth. |
9–24, brown to yellow, ellipsoid, ovoid, or obovoid, flat, 0.8–1.6 × (0.3–)0.6–1.2 mm, 0.2–0.3 mm thick, smooth. |
20–30, brownish, ellipsoid to ovoid, indistinct-compressed or planoconvex, 0.5–0.7 × 0.3–0.6 mm, thickness varies due to compression in capsule, indistinctly wrinkled or ± rugose. |
2n | = 52. |
= 18, 36 (Europe). |
= 36. |
Veronica peregrina |
Veronica officinalis |
Veronica americana |
|
Phenology | Flowering (Feb–)Mar–Jun(–Nov). | Flowering (Apr–)Jun–Jul(–Aug). | Flowering May–Aug. |
Habitat | Moist waste lands, gardens, roadsides, stream banks, pond shorelines, vernal pools, other cultivated land. | Forests, meadows, pastures, scrub, ruderal places. | Slowly flowing waters, banks, sand bars, gravel flood plains, moist soils, springs, ditches, swamps, marshes. |
Elevation | 0–700(–3000) m. [0–2300(–9800) ft.] | 0–1600 m. [0–5200 ft.] | 0–3600 m. [0–11800 ft.] |
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Central America; South America; Mexico (Baja California, Baja California Sur, Chihuahua, México, Sonora); Eurasia; Australia
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CA; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Eurasia [Introduced in North America; introduced in s South America (Chile)]
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AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Mexico; Asia (Japan, Russia)
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Discussion | Stem indument of Veronica peregrina has been used to distinguish var. xalapensis with a glandular-hairy stem (F. W. Pennell 1935), a variety that seems to be confined to drier places than the type variety. Variety laurentiana is also differentiated from var. peregrina in having a glandular-hairy stem, fleshier leaves, shorter stamens, smaller corollas, and slightly differently shaped capsules (Frère Marie-Victorin and J. Rousseau 1940). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica ×tournefortii Villars [V. officinalis var. tournefortii (Villars) Dumortier] refers to the hybrid between V. officinalis and V. allionii Villars in Europe and is not applicable to any North American plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although difficult to separate, Veronica beccabunga and V. americana are maintained as species here. Evidence suggesting separate species status is different ploidy level (the tetraploid level is rare in V. beccabunga) and the occurrence of V. americana in more natural habitats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 314. | FNA vol. 17, p. 311. | FNA vol. 17, p. 313. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | |||
Synonyms | V. peregrina var. laurentiana, V. peregrina subsp. xalapensis, V. peregrina var. xalapensis, V. sherwoodii, V. xalapensis | V. beccabunga var. americana | |
Name authority | Linnaeus: Sp. Pl. 1: 14. (1753) | Linnaeus: Sp. Pl. 1: 11. (1753) | Schweinitz ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 468. (1846) |
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