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common gypsyweed, common speedwell, heath speedwell, herbal speedwell, Paul's betony, véronique officinale

Cusick's speedwell

Habit Perennials. Perennials.
Stems

creeping, decumbent to ascending distally, 10–40(–50) cm, densely eglandular-hairy, rarely glabrate or glabrous.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade ovate, ovate-oblong, or obovate, (8–)20–35(–50) × (3–)10–20(–30) mm, 1.5–2 times as long as wide, base attenuate to cuneate or obtuse, margins dentate, denticulate, or serrulate, apex obtuse to rounded, surfaces sparsely to densely eglandular-hairy, rarely glabrate.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1–4(–8), axillary, 40–60 mm, 10–35(–40)-flowered, axis densely to sparsely eglandular-hairy, sometimes also glandular-hairy, rarely glabrate;

bracts linear-lanceolate or narrowly ovate, (1.5–)3–4(–5) mm.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

erect, (0.5–)1–2(–3) mm, shorter than subtending bract, densely eglandular-hairy, rarely glandular-hairy or glabrate.

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx 4(or 5)-lobed, lobes 2–3.5(–4) mm, apex obtuse, glandular- and eglandular-hairy;

corolla pale blue-lilac to pale pink, rarely white with pink veins, 6–8 mm diam.;

stamens 2.8–3.4 mm;

style 2.5–3(–4) mm.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

compressed in cross section, triangulate-obdeltoid to obcordiform, 3–5 × 4–5 mm, apex rounded or truncate, rarely emarginate, glandular-hairy, rarely glabrous.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

9–24, brown to yellow, ellipsoid, ovoid, or obovoid, flat, 0.8–1.6 × (0.3–)0.6–1.2 mm, 0.2–0.3 mm thick, smooth.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 18, 36 (Europe).

= 18, 72.

Veronica officinalis

Veronica cusickii

Phenology Flowering (Apr–)Jun–Jul(–Aug). Flowering Jun–Aug.
Habitat Forests, meadows, pastures, scrub, ruderal places. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 0–1600 m. (0–5200 ft.) 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
CA; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Eurasia [Introduced in North America; introduced in s South America (Chile)]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica ×tournefortii Villars [V. officinalis var. tournefortii (Villars) Dumortier] refers to the hybrid between V. officinalis and V. allionii Villars in Europe and is not applicable to any North American plant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 311. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority Linnaeus: Sp. Pl. 1: 11. (1753) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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