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ivy speedwell, ivy-leaf speedwell

Cusick's speedwell

Habit Annuals. Perennials.
Stems

decumbent to ascending, 5–40(–50) cm, eglandular-hairy.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade suborbiculate, (3–)5-lobed, central lobe usually overtopping lateral ones, (5–)7–15(–20) × 8–16(–25) mm, base truncate, margins (3–)5-lobed, apex acute, surfaces sparsely eglandular-hairy.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1–10, terminal, 50–400(–500) mm, 5–20-flowered, axis eglandular-hairy;

bracts suborbiculate, (3–)5-lobed, central lobe usually overtopping lateral ones, (5–)7–15(–20) mm.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

patent or deflexed, (5–)9–15(–20) mm, equal to or ± shorter than subtending bract, length 2–3 times calyx, eglandular-hairy (in single line adaxially).

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes 5–6(–7) mm, apex acute, abaxial surface usually glabrous, ciliate with 25–35 hairs per side;

corolla bright blue with bright white center, 5–7(–9) mm diam.;

stamens sky blue, 0.7–1.2 mm;

style (0.6–)0.7–0.9(–1.1) mm.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

± round in cross section, ovoid, 3–4 × 4.5–6 mm, apex ± emarginate, glabrate.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

1–4, bright yellow, ellipsoid to subglobular, urn-shaped, 2.3–3.3 × 2–3 mm, 1–2.2 mm thick, weakly cristate or rugose, ± smooth.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 54 (Eurasia).

= 18, 72.

Veronica hederifolia

Veronica cusickii

Phenology Flowering (Feb–)Mar–Jun. Flowering Jun–Aug.
Habitat Fields, lawns, gardens, ruderal places, vineyards, open forests, shady rocky places, dunes. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 0–2000 m. (0–6600 ft.) 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MI; MO; NC; NE; NJ; NY; OH; OK; OR; PA; SC; SD; TN; UT; VA; WA; WV; BC; ON; Eurasia [Introduced in North America]
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from FNA
CA; ID; MT; OR; WA; BC
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[BONAP county map]
Discussion

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 315. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority Linnaeus: Sp. Pl. 1: 13. (1753) — (as hederaefolia) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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