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rock speedwell, woodystem speedwell

Copeland's speedwell

Habit Perennials. Perennials.
Stems

ascending to erect, densely branched from woody base, (5–)10–15(–30) cm, eglandular-hairy, sometimes glandular-hairy.

ascending, unbranched, 5–15 cm, densely glandular-hairy.

Leaves

blade obovate to ovate or spatulate, sometimes suborbiculate proximally, (7–)8–20(–25) × (2–)3–6(–7) mm, mostly shorter to equal to internodes, base cuneate, margins entire or ± crenate or serrate, apex acute, surfaces glabrous or glabrate.

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

Racemes

1–10(–20), terminal, rarely with 1–4 axillary, 20–40 mm, (1–)4–10(–18)-flowered, axis glabrate to sparsely eglandular-hairy;

bracts linear or linear-lanceolate or long-ovate, (1–)3.5–9(–15) mm.

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

Pedicels

erect, (2–)5–7(–15) mm, equal to ± longer than subtending bracts, eglandular-hairy.

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

Flowers

calyx 4(or 5)-lobed, lobes (2.5–)4.5–6(–8) mm, apex obtuse, eglandular-hairy, sometimes also glandular-hairy;

corolla intense blue, sometimes with reddish or white center, rarely white, rotate, (6–)10–14(–15) mm diam.;

stamens 5–7 mm;

style (3–)5–7(–8.5) mm.

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

Capsules

± compressed in cross section, ovoid, (5–)6–8(–9) × (3–)4–4.5(–5.5) mm, longer than wide, apex attenuate to apiculate and acute, rarely rounded, not emarginate, eglandular-hairy, sometimes mixed with larger glandular hairs.

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

Seeds

35–62, orange, ellipsoid, flat, 0.9–1.5 × (0.4–)0.9–1.2 mm, 0.2–0.3 mm thick, smooth.

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

2n

= 16.

= 18.

Veronica fruticans

Veronica copelandii

Phenology Flowering (May–)Jun–Aug. Flowering Jul–Aug.
Habitat Fissures, rocky places, scree. Subalpine meadows, alpine slopes.
Elevation 0–1000 m. (0–3300 ft.) 1900–2500 m. (6200–8200 ft.)
Distribution
from USDA
Greenland
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 318. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Jacquin: Enum. Stirp. Vindob. 2: 200. (1762) Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi)
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