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rock speedwell, woodystem speedwell

bilobed speedwell, twolobe speedwell

Habit Perennials. Annuals.
Stems

ascending to erect, densely branched from woody base, (5–)10–15(–30) cm, eglandular-hairy, sometimes glandular-hairy.

erect, (1–)2–10(–18) cm, eglandular-hairy.

Leaves

blade obovate to ovate or spatulate, sometimes suborbiculate proximally, (7–)8–20(–25) × (2–)3–6(–7) mm, mostly shorter to equal to internodes, base cuneate, margins entire or ± crenate or serrate, apex acute, surfaces glabrous or glabrate.

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

Racemes

1–10(–20), terminal, rarely with 1–4 axillary, 20–40 mm, (1–)4–10(–18)-flowered, axis glabrate to sparsely eglandular-hairy;

bracts linear or linear-lanceolate or long-ovate, (1–)3.5–9(–15) mm.

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

Pedicels

erect, (2–)5–7(–15) mm, equal to ± longer than subtending bracts, eglandular-hairy.

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

Flowers

calyx 4(or 5)-lobed, lobes (2.5–)4.5–6(–8) mm, apex obtuse, eglandular-hairy, sometimes also glandular-hairy;

corolla intense blue, sometimes with reddish or white center, rarely white, rotate, (6–)10–14(–15) mm diam.;

stamens 5–7 mm;

style (3–)5–7(–8.5) mm.

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

Capsules

± compressed in cross section, ovoid, (5–)6–8(–9) × (3–)4–4.5(–5.5) mm, longer than wide, apex attenuate to apiculate and acute, rarely rounded, not emarginate, eglandular-hairy, sometimes mixed with larger glandular hairs.

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

Seeds

35–62, orange, ellipsoid, flat, 0.9–1.5 × (0.4–)0.9–1.2 mm, 0.2–0.3 mm thick, smooth.

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

2n

= 16.

= 28 (Asia).

Veronica fruticans

Veronica biloba

Phenology Flowering (May–)Jun–Aug. Flowering Apr–Jul.
Habitat Fissures, rocky places, scree. Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils.
Elevation 0–1000 m. (0–3300 ft.) (400–)1300–3000 m. ((1300–)4300–9800 ft.)
Distribution
from USDA
Greenland
[BONAP county map]
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 318. FNA vol. 17, p. 321.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Jacquin: Enum. Stirp. Vindob. 2: 200. (1762) Linnaeus: Mant. Pl. 2: 172. (1771)
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