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Dillenius' speedwell

Cusick's speedwell

Habit Annuals. Perennials.
Stems

erect to ascending, (8–)10–20(–40) cm, glandular- and eglandular-hairy.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade lanceolate to ovate, 7–19(–21) × (3–)5–12 mm, 1–2 times as long as wide, base cuneate, margins (proximal) crenate-dentate or (distal) ±palmatifid, lobes 3–7+, apex of central lobe obtuse, lateral ± acute, surfaces glandular- and eglandular-hairy.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1, terminal, 60–180 mm, 15–50(–120)-flowered, axis eglandular- and glandular-hairy;

proximal bracts similar to leaves, distal ones linear-lanceolate, 3–5 mm.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

erect, 2–4(–5) mm, shorter than subtending bract, length 1/2–1 times calyx, eglandular- and glandular-hairy.

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes 3.5–6 mm, apex acute, eglandular- and glandular-hairy;

corolla deep blue, 4–6 mm diam.;

stamens 0.8–1.5 mm;

style 0.8–1.5 mm, stigma violet.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

compressed in cross section, narrowly obcordiform, 3.5–4.5 × 4–6 mm, apex emarginate, angle of sinus ca. 90°, ciliate with glandular and eglandular hairs.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

10–28, yellowish, ellipsoid, flat, 0.9–1.6 × 0.7–1.3 mm, 0.2–0.3 mm thick, smooth.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 16 (Eurasia).

= 18, 72.

Veronica dillenii

Veronica cusickii

Phenology Flowering (Mar–)Apr–Jul. Flowering Jun–Aug.
Habitat Open pine and oak forests, rocky, dry, sandy slopes. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 500–2200 m. (1600–7200 ft.) 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
IL; IN; MI; NY; VA; WI; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica dillenii is closely related to V. verna but with larger flowers; it may have been overlooked and may be distributed more widely. Most herbarium specimens of V. dillenii blacken when dry due to the presence of aucubin, which distinguishes them from V. verna.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 318. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority Crantz: Stirp. Austr. Fasc. ed. 2, 2: 352. (1769) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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