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Dillenius' speedwell

bilobed speedwell, sawtooth speedwell, twolobed speedwell

Habit Annuals. Annuals.
Stems

erect to ascending, (8–)10–20(–40) cm, glandular- and eglandular-hairy.

erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy.

Leaves

blade lanceolate to ovate, 7–19(–21) × (3–)5–12 mm, 1–2 times as long as wide, base cuneate, margins (proximal) crenate-dentate or (distal) ±palmatifid, lobes 3–7+, apex of central lobe obtuse, lateral ± acute, surfaces glandular- and eglandular-hairy.

blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate.

Racemes

1, terminal, 60–180 mm, 15–50(–120)-flowered, axis eglandular- and glandular-hairy;

proximal bracts similar to leaves, distal ones linear-lanceolate, 3–5 mm.

1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy;

bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire.

Pedicels

erect, 2–4(–5) mm, shorter than subtending bract, length 1/2–1 times calyx, eglandular- and glandular-hairy.

erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy.

Flowers

calyx lobes 3.5–6 mm, apex acute, eglandular- and glandular-hairy;

corolla deep blue, 4–6 mm diam.;

stamens 0.8–1.5 mm;

style 0.8–1.5 mm, stigma violet.

calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy;

corolla mostly intense blue with white center, 4–6 mm diam.;

stamens 2.5 mm;

style (0.7–)1–1.3(–1.7) mm.

Capsules

compressed in cross section, narrowly obcordiform, 3.5–4.5 × 4–6 mm, apex emarginate, angle of sinus ca. 90°, ciliate with glandular and eglandular hairs.

slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy.

Seeds

10–28, yellowish, ellipsoid, flat, 0.9–1.6 × 0.7–1.3 mm, 0.2–0.3 mm thick, smooth.

4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate.

2n

= 16 (Eurasia).

= 18, 42 (Asia).

Veronica dillenii

Veronica argute-serrata

Phenology Flowering (Mar–)Apr–Jul. Flowering Apr–Jun.
Habitat Open pine and oak forests, rocky, dry, sandy slopes. Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils.
Elevation 500–2200 m. (1600–7200 ft.) (600–)900–3000 m. ((2000–)3000–9800 ft.)
Distribution
from FNA
IL; IN; MI; NY; VA; WI; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
[WildflowerSearch map]
Discussion

Veronica dillenii is closely related to V. verna but with larger flowers; it may have been overlooked and may be distributed more widely. Most herbarium specimens of V. dillenii blacken when dry due to the presence of aucubin, which distinguishes them from V. verna.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 318. FNA vol. 17, p. 321.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. bornmuelleri
Name authority Crantz: Stirp. Austr. Fasc. ed. 2, 2: 352. (1769) Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata)
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