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glandular speedwell, pale speedwell

common gypsyweed, common speedwell, heath speedwell, herbal speedwell, Paul's betony, véronique officinale

Habit Annuals. Perennials.
Stems

decumbent to ascending, (5–)10–30 cm, sparsely subvillous to glabrate.

creeping, decumbent to ascending distally, 10–40(–50) cm, densely eglandular-hairy, rarely glabrate or glabrous.

Leaves

blade suborbiculate, 6–20 × 9–20 mm, base subcordate, truncate, or cuneate, margins 5–9-lobed 1/6–1/3 of width of blade, lateral lobes ± narrower, apex acute, surfaces eglandular-hairy, rarely also glandular-hairy or glabrate.

blade ovate, ovate-oblong, or obovate, (8–)20–35(–50) × (3–)10–20(–30) mm, 1.5–2 times as long as wide, base attenuate to cuneate or obtuse, margins dentate, denticulate, or serrulate, apex obtuse to rounded, surfaces sparsely to densely eglandular-hairy, rarely glabrate.

Racemes

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis eglandular-hairy, rarely also glandular-hairy or glabrate;

bracts suborbiculate, 6–20 mm, 5- to 9-lobed for 1/6–1/3 of width of leaf blade, lateral lobes ± narrower.

1–4(–8), axillary, 40–60 mm, 10–35(–40)-flowered, axis densely to sparsely eglandular-hairy, sometimes also glandular-hairy, rarely glabrate;

bracts linear-lanceolate or narrowly ovate, (1.5–)3–4(–5) mm.

Pedicels

patent or recurved, 5–15(–30) mm, equal in flower to ± longer than subtending bract in fruit, eglandular- or glandular-pilose or glabrous.

erect, (0.5–)1–2(–3) mm, shorter than subtending bract, densely eglandular-hairy, rarely glandular-hairy or glabrate.

Flowers

calyx lobes erect-patent, (3–)4–5 mm, apex obtuse, abaxial surfaces eglandular- or glandular-hairy, rarely glabrous, not ciliate;

corolla white, 6–12 mm diam.;

stamens 1–2 mm;

style 1–2 mm.

calyx 4(or 5)-lobed, lobes 2–3.5(–4) mm, apex obtuse, glandular- and eglandular-hairy;

corolla pale blue-lilac to pale pink, rarely white with pink veins, 6–8 mm diam.;

stamens 2.8–3.4 mm;

style 2.5–3(–4) mm.

Capsules

± round in cross section, ovoid to ellipsoid, 2.5–3.5 × 3.5–4.5 mm, apex ± emarginate, eglandular- or glandular-pilose.

compressed in cross section, triangulate-obdeltoid to obcordiform, 3–5 × 4–5 mm, apex rounded or truncate, rarely emarginate, glandular-hairy, rarely glabrous.

Seeds

1–4, brownish orange, subglobular, urn-shaped (elaiosome inside), (2–)2.5–3.1 × (1.7–)2–2.5(–2.8) mm, 1.1–1.7 mm thick, cristate.

9–24, brown to yellow, ellipsoid, ovoid, or obovoid, flat, 0.8–1.6 × (0.3–)0.6–1.2 mm, 0.2–0.3 mm thick, smooth.

2n

= 36, 54 (Eurasia).

= 18, 36 (Europe).

Veronica cymbalaria

Veronica officinalis

Phenology Flowering Mar–May(–Jun). Flowering (Apr–)Jun–Jul(–Aug).
Habitat Lawns. Forests, meadows, pastures, scrub, ruderal places.
Elevation 20 m. (100 ft.) 0–1600 m. (0–5200 ft.)
Distribution
from FNA
LA; Eurasia (Mediterranean) [Introduced in North America]
[BONAP county map]
from FNA
CA; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Eurasia [Introduced in North America; introduced in s South America (Chile)]
[WildflowerSearch map]
[BONAP county map]
Discussion

The occurrence of Veronica cymbalaria in other states is possible. It has been reported from Maryland; a herbarium specimen could not be located. The report seems to be based on incorrect synonymy (C. Frye, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica ×tournefortii Villars [V. officinalis var. tournefortii (Villars) Dumortier] refers to the hybrid between V. officinalis and V. allionii Villars in Europe and is not applicable to any North American plant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 311.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Bodard: Mém. Véronique Cymb., 3. (1798) Linnaeus: Sp. Pl. 1: 11. (1753)
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