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glandular speedwell, pale speedwell

Cusick's speedwell

Habit Annuals. Perennials.
Stems

decumbent to ascending, (5–)10–30 cm, sparsely subvillous to glabrate.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade suborbiculate, 6–20 × 9–20 mm, base subcordate, truncate, or cuneate, margins 5–9-lobed 1/6–1/3 of width of blade, lateral lobes ± narrower, apex acute, surfaces eglandular-hairy, rarely also glandular-hairy or glabrate.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis eglandular-hairy, rarely also glandular-hairy or glabrate;

bracts suborbiculate, 6–20 mm, 5- to 9-lobed for 1/6–1/3 of width of leaf blade, lateral lobes ± narrower.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

patent or recurved, 5–15(–30) mm, equal in flower to ± longer than subtending bract in fruit, eglandular- or glandular-pilose or glabrous.

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes erect-patent, (3–)4–5 mm, apex obtuse, abaxial surfaces eglandular- or glandular-hairy, rarely glabrous, not ciliate;

corolla white, 6–12 mm diam.;

stamens 1–2 mm;

style 1–2 mm.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

± round in cross section, ovoid to ellipsoid, 2.5–3.5 × 3.5–4.5 mm, apex ± emarginate, eglandular- or glandular-pilose.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

1–4, brownish orange, subglobular, urn-shaped (elaiosome inside), (2–)2.5–3.1 × (1.7–)2–2.5(–2.8) mm, 1.1–1.7 mm thick, cristate.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 36, 54 (Eurasia).

= 18, 72.

Veronica cymbalaria

Veronica cusickii

Phenology Flowering Mar–May(–Jun). Flowering Jun–Aug.
Habitat Lawns. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 20 m. (100 ft.) 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
LA; Eurasia (Mediterranean) [Introduced in North America]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

The occurrence of Veronica cymbalaria in other states is possible. It has been reported from Maryland; a herbarium specimen could not be located. The report seems to be based on incorrect synonymy (C. Frye, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority Bodard: Mém. Véronique Cymb., 3. (1798) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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