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glandular speedwell, pale speedwell

Copeland's speedwell

Habit Annuals. Perennials.
Stems

decumbent to ascending, (5–)10–30 cm, sparsely subvillous to glabrate.

ascending, unbranched, 5–15 cm, densely glandular-hairy.

Leaves

blade suborbiculate, 6–20 × 9–20 mm, base subcordate, truncate, or cuneate, margins 5–9-lobed 1/6–1/3 of width of blade, lateral lobes ± narrower, apex acute, surfaces eglandular-hairy, rarely also glandular-hairy or glabrate.

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

Racemes

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis eglandular-hairy, rarely also glandular-hairy or glabrate;

bracts suborbiculate, 6–20 mm, 5- to 9-lobed for 1/6–1/3 of width of leaf blade, lateral lobes ± narrower.

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

Pedicels

patent or recurved, 5–15(–30) mm, equal in flower to ± longer than subtending bract in fruit, eglandular- or glandular-pilose or glabrous.

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

Flowers

calyx lobes erect-patent, (3–)4–5 mm, apex obtuse, abaxial surfaces eglandular- or glandular-hairy, rarely glabrous, not ciliate;

corolla white, 6–12 mm diam.;

stamens 1–2 mm;

style 1–2 mm.

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

Capsules

± round in cross section, ovoid to ellipsoid, 2.5–3.5 × 3.5–4.5 mm, apex ± emarginate, eglandular- or glandular-pilose.

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

Seeds

1–4, brownish orange, subglobular, urn-shaped (elaiosome inside), (2–)2.5–3.1 × (1.7–)2–2.5(–2.8) mm, 1.1–1.7 mm thick, cristate.

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

2n

= 36, 54 (Eurasia).

= 18.

Veronica cymbalaria

Veronica copelandii

Phenology Flowering Mar–May(–Jun). Flowering Jul–Aug.
Habitat Lawns. Subalpine meadows, alpine slopes.
Elevation 20 m. (100 ft.) 1900–2500 m. (6200–8200 ft.)
Distribution
from FNA
LA; Eurasia (Mediterranean) [Introduced in North America]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

The occurrence of Veronica cymbalaria in other states is possible. It has been reported from Maryland; a herbarium specimen could not be located. The report seems to be based on incorrect synonymy (C. Frye, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Bodard: Mém. Véronique Cymb., 3. (1798) Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi)
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