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glandular speedwell, pale speedwell

bird's-eye speedwell, Germander speedwell, véronique petit-chêne

Habit Annuals. Perennials.
Stems

decumbent to ascending, (5–)10–30 cm, sparsely subvillous to glabrate.

ascending, rarely erect, (7–)10–30(–50) cm, densely eglandular-hairy, hairs along stem in 2 prominent lines.

Leaves

blade suborbiculate, 6–20 × 9–20 mm, base subcordate, truncate, or cuneate, margins 5–9-lobed 1/6–1/3 of width of blade, lateral lobes ± narrower, apex acute, surfaces eglandular-hairy, rarely also glandular-hairy or glabrate.

blade narrowly ovate to ovate-orbiculate, (10–)12–30(–42) × (6–)10–22(–30) mm, 1–2 times as long as wide, base truncate to ± cordate, margins crenate to deeply incised, apex obtuse, surfaces variably hairy.

Racemes

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis eglandular-hairy, rarely also glandular-hairy or glabrate;

bracts suborbiculate, 6–20 mm, 5- to 9-lobed for 1/6–1/3 of width of leaf blade, lateral lobes ± narrower.

1–4, axillary, 40–100(–200) mm, 15–40(–60)-flowered, axis eglandular-hairy, sometimes also glandular-hairy;

bracts linear-elliptic, 3–7 mm.

Pedicels

patent or recurved, 5–15(–30) mm, equal in flower to ± longer than subtending bract in fruit, eglandular- or glandular-pilose or glabrous.

suberect, (3–)5–8(–10) mm, equal to or shorter than subtending bract, eglandular- and glandular-hairy.

Flowers

calyx lobes erect-patent, (3–)4–5 mm, apex obtuse, abaxial surfaces eglandular- or glandular-hairy, rarely glabrous, not ciliate;

corolla white, 6–12 mm diam.;

stamens 1–2 mm;

style 1–2 mm.

calyx 4-lobed, lobes 2–8 mm, apex acute, eglandular- and, sometimes, glandular-hairy;

corolla blue obscure darker nerves and sometimes whitish margin, (6–)10–14(–17) mm diam.;

stamens 4.5–6.5 mm;

style (2.5–)4–5 mm.

Capsules

± round in cross section, ovoid to ellipsoid, 2.5–3.5 × 3.5–4.5 mm, apex ± emarginate, eglandular- or glandular-pilose.

strongly compressed in cross section, obcordiform to obdeltoid, (2–)3.5–4(–5) × (3.5–)4–5(–5.5) mm, apex ± emarginate, eglandular-hairy.

Seeds

1–4, brownish orange, subglobular, urn-shaped (elaiosome inside), (2–)2.5–3.1 × (1.7–)2–2.5(–2.8) mm, 1.1–1.7 mm thick, cristate.

(2–)12–20(–28), yellow, ellipsoid, flat, 1.1–1.7 × 0.6–1.5 mm, 0.2–0.4 mm thick, smooth to subrugose.

2n

= 36, 54 (Eurasia).

= 16, 32 (Eurasia).

Veronica cymbalaria

Veronica chamaedrys

Phenology Flowering Mar–May(–Jun). Flowering Apr–Jun(–Oct).
Habitat Lawns. Rich soils, deciduous forests, forest edges, roadsides, chaparral, scrub, meadows, lawns.
Elevation 20 m. (100 ft.) 0–2200 m. (0–7200 ft.)
Distribution
from FNA
LA; Eurasia (Mediterranean) [Introduced in North America]
[BONAP county map]
from FNA
AK; CT; DC; ID; IL; IN; MA; MD; ME; MI; MO; MT; NC; NH; NJ; NY; OH; OR; PA; RI; VA; VT; WA; WI; WV; AB; BC; NB; NF; NS; ON; PE; QC; Eurasia [Introduced in South America (Argentina)]
[WildflowerSearch map]
[BONAP county map]
Discussion

The occurrence of Veronica cymbalaria in other states is possible. It has been reported from Maryland; a herbarium specimen could not be located. The report seems to be based on incorrect synonymy (C. Frye, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The description provided here for Veronica chamaedrys is solely for the tetraploid cytotype, most probably the exclusive cytotype in the flora area and in central and western Europe. The diploid cytotype is so far only known from eastern and southeastern Europe (K. E. Bardy et al. 2010). A significant change in morphology can occur in shaded habitats, in which especially the petiole can be elongated beyond the range given.

It is unclear whether Veronica chamaedrys is introduced throughout the flora area; it may be native in northeastern areas of North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 318.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Bodard: Mém. Véronique Cymb., 3. (1798) Linnaeus: Sp. Pl. 1: 13. (1753)
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