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glandular speedwell, pale speedwell

American brooklime, American speedwell, véronique d'amérique

Habit Annuals. Perennials.
Stems

decumbent to ascending, (5–)10–30 cm, sparsely subvillous to glabrate.

decumbent or ascending, 5–50 cm, glabrous.

Leaves

blade suborbiculate, 6–20 × 9–20 mm, base subcordate, truncate, or cuneate, margins 5–9-lobed 1/6–1/3 of width of blade, lateral lobes ± narrower, apex acute, surfaces eglandular-hairy, rarely also glandular-hairy or glabrate.

petiole 2–6(–10) mm;

blade ovate to lanceolate, widest proximal to middle, (5–)30–50(–100) × (3–)7–20(–30) mm, 2–4 times as long as wide, base truncate, rounded or almost cordate, abruptly turning into petioles, margins entire or serrate, apex obtuse-orbicular or acute, surfaces glabrous.

Racemes

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis eglandular-hairy, rarely also glandular-hairy or glabrate;

bracts suborbiculate, 6–20 mm, 5- to 9-lobed for 1/6–1/3 of width of leaf blade, lateral lobes ± narrower.

1–8, axillary, 30–100 mm, (3–)10–30-flowered, axis glabrous;

bracts lanceolate, 3–6 mm.

Pedicels

patent or recurved, 5–15(–30) mm, equal in flower to ± longer than subtending bract in fruit, eglandular- or glandular-pilose or glabrous.

patent, 3–10(–12) mm, equal to ± longer than subtending bract, glabrous.

Flowers

calyx lobes erect-patent, (3–)4–5 mm, apex obtuse, abaxial surfaces eglandular- or glandular-hairy, rarely glabrous, not ciliate;

corolla white, 6–12 mm diam.;

stamens 1–2 mm;

style 1–2 mm.

calyx lobes 2–5(–6) mm, apex acute, glabrous;

corolla pink or sky blue, 4–10 mm diam.;

stamens 2–4 mm;

style (1.7–)2.5–3.5(–4) mm.

Capsules

± round in cross section, ovoid to ellipsoid, 2.5–3.5 × 3.5–4.5 mm, apex ± emarginate, eglandular- or glandular-pilose.

slightly compressed in cross section, globular, 2–4 × 3–5 mm, apex not or ± emarginate, glabrous.

Seeds

1–4, brownish orange, subglobular, urn-shaped (elaiosome inside), (2–)2.5–3.1 × (1.7–)2–2.5(–2.8) mm, 1.1–1.7 mm thick, cristate.

20–30, brownish, ellipsoid to ovoid, indistinct-compressed or planoconvex, 0.5–0.7 × 0.3–0.6 mm, thickness varies due to compression in capsule, indistinctly wrinkled or ± rugose.

2n

= 36, 54 (Eurasia).

= 36.

Veronica cymbalaria

Veronica americana

Phenology Flowering Mar–May(–Jun). Flowering May–Aug.
Habitat Lawns. Slowly flowing waters, banks, sand bars, gravel flood plains, moist soils, springs, ditches, swamps, marshes.
Elevation 20 m. (100 ft.) 0–3600 m. (0–11800 ft.)
Distribution
from FNA
LA; Eurasia (Mediterranean) [Introduced in North America]
[BONAP county map]
from FNA
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Mexico; Asia (Japan, Russia)
[WildflowerSearch map]
[BONAP county map]
Discussion

The occurrence of Veronica cymbalaria in other states is possible. It has been reported from Maryland; a herbarium specimen could not be located. The report seems to be based on incorrect synonymy (C. Frye, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although difficult to separate, Veronica beccabunga and V. americana are maintained as species here. Evidence suggesting separate species status is different ploidy level (the tetraploid level is rare in V. beccabunga) and the occurrence of V. americana in more natural habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 313.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. beccabunga var. americana
Name authority Bodard: Mém. Véronique Cymb., 3. (1798) Schweinitz ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 468. (1846)
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