Veronica cusickii |
Veronica wormskjoldii |
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Cusick's speedwell |
alpine speedwell, American alpine speedwell, hairy speedwell, spring speedwell, thyme-leaf speedwell, wormskjold's alpine speedwell |
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Habit | Perennials. | Perennials. | ||||
Stems | erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy. |
erect or ascending, light grayish green distally, unbranched, (3–)8–50 cm, sparsely to densely villous-hirsute, sometimes also glandular-hairy. |
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Leaves | blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate. |
blade elliptic to lanceolate or oblong-ovate, 8–40 × 5–20 mm, base cuneate, margins entire, dentate, or serrate, apex short-acuminate, surfaces sparsely to densely villous-hirsute or glabrous. |
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Racemes | 1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy; bracts lanceolate, 3–7(–11) mm. |
1, terminal, 5–40 mm, to 60(–150) mm in fruit, (2–)5–25-flowered, axis densely villous-hirsute and ± glandular-hairy; bracts linear to lanceolate, 1–8 mm. |
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Pedicels | erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy. |
erect, 2–10(–15) mm, ca. equal to subtending bract, densely villous-hirsute or glandular-hairy. |
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Flowers | calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy; corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.; stamens 4–8 mm; style (5–)6–9(–10) mm, conspicuously exerted. |
calyx lobes (2.5–)3–5.5(–7) mm, apex obtuse or acuminate, glabrous or densely glandular-hairy; corolla deep blue-violet or violet-purple to deep blue, rotate, 3–11 mm diam.; stamens 1–2 mm; style 0.8–4(–6) mm. |
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Capsules | compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy. |
compressed in cross section, oblong-obovoid, 4–6(–8) × (2.8–)4–5.5 mm, ca. as long as wide, apex emarginate, usually densely glandular-hairy. |
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Seeds | 28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough. |
10–50, straw colored, ellipsoid, flat, 0.7–1 × 0.4–1 mm, 0.05–1 mm thick, very minutely striate. |
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2n | = 18, 72. |
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Veronica cusickii |
Veronica wormskjoldii |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows. | |||||
Elevation | 1500–3200 m. (4900–10500 ft.) | |||||
Distribution |
CA; ID; MT; OR; WA; BC
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AK; AZ; CA; CO; ID; ME; MT; NH; NM; NV; OR; UT; WA; WY; AB; BC; NT; NU; ON; QC; YT
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Discussion | Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Veronica wormskjoldii occurs in two cytotypes, a diploid (2n = 18) in western North America and a tetraploid (2n = 36) in eastern North America and Greenland. As demonstrated by D. C. Albach et al. (2006), the tetraploid is likely to be a hybrid of the diploid V. alpina and V. wormskjoldii. In that publication, species rank was used for simplicity, not as a taxonomic conclusion. However, subsequent morphological analyses in conjunction with preparing this treatment revealed that the large variation in the diploid V. wormskjoldii, also supported by the large number of varieties established within that taxon, make it nearly impossible to differentiate the two North American taxa morphologically. In cases where two taxa are morphologically undifferentiable, but karyologically and genetically distinct and geographically clearly separate, the rank of subspecies seems more appropriate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 310. | FNA vol. 17, p. 309. | ||||
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. allenii | V. alpina subsp. wormskjoldii, V. alpina var. wormskjoldii | ||||
Name authority | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878) | Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 1: 101. (1817) — (as wormskjoldi) | ||||
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