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Cusick's speedwell

Dillenius' speedwell

Habit Perennials. Annuals.
Stems

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

erect to ascending, (8–)10–20(–40) cm, glandular- and eglandular-hairy.

Leaves

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

blade lanceolate to ovate, 7–19(–21) × (3–)5–12 mm, 1–2 times as long as wide, base cuneate, margins (proximal) crenate-dentate or (distal) ±palmatifid, lobes 3–7+, apex of central lobe obtuse, lateral ± acute, surfaces glandular- and eglandular-hairy.

Racemes

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

1, terminal, 60–180 mm, 15–50(–120)-flowered, axis eglandular- and glandular-hairy;

proximal bracts similar to leaves, distal ones linear-lanceolate, 3–5 mm.

Pedicels

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

erect, 2–4(–5) mm, shorter than subtending bract, length 1/2–1 times calyx, eglandular- and glandular-hairy.

Flowers

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

calyx lobes 3.5–6 mm, apex acute, eglandular- and glandular-hairy;

corolla deep blue, 4–6 mm diam.;

stamens 0.8–1.5 mm;

style 0.8–1.5 mm, stigma violet.

Capsules

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

compressed in cross section, narrowly obcordiform, 3.5–4.5 × 4–6 mm, apex emarginate, angle of sinus ca. 90°, ciliate with glandular and eglandular hairs.

Seeds

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

10–28, yellowish, ellipsoid, flat, 0.9–1.6 × 0.7–1.3 mm, 0.2–0.3 mm thick, smooth.

2n

= 18, 72.

= 16 (Eurasia).

Veronica cusickii

Veronica dillenii

Phenology Flowering Jun–Aug. Flowering (Mar–)Apr–Jul.
Habitat Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows. Open pine and oak forests, rocky, dry, sandy slopes.
Elevation 1500–3200 m. (4900–10500 ft.) 500–2200 m. (1600–7200 ft.)
Distribution
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
IL; IN; MI; NY; VA; WI; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica dillenii is closely related to V. verna but with larger flowers; it may have been overlooked and may be distributed more widely. Most herbarium specimens of V. dillenii blacken when dry due to the presence of aucubin, which distinguishes them from V. verna.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 310. FNA vol. 17, p. 318.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878) Crantz: Stirp. Austr. Fasc. ed. 2, 2: 352. (1769)
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