The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Copeland's speedwell

undulate speedwell, wavy-leaf water speedwell

Habit Perennials. Annuals or perennials.
Stems

ascending, unbranched, 5–15 cm, densely glandular-hairy.

erect or prostrate basally, 10–100 cm, glabrous at least proximally, glandular-hairy distally.

Leaves

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

petiole 0–5 mm;

blade elliptic to ovate, sometimes ovate-oblong or linear-lanceolate, rarely lanceolate, 20–50(–100) × 5–20(–25) mm, 2.5–4 times as long as wide, base attenuate, upwards amplexicaul, margins subentire or crenate to serrate or ± undulate, apex acute to obtuse, surfaces glabrous.

Racemes

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

6–25, axillary, 50–220 mm, 10–15 mm diam., 10–100-flowered, axis sparsely glandular-hairy, rarely glabrous;

pedicels 2–6 per cm;

bracts linear to lanceolate, 3–6 mm, apex acute.

Pedicels

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

patent, 3–5 mm, equal to subtending bract, sparsely glandular-hairy.

Flowers

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

calyx lobes 1.5–2(–3) mm, apex acute, sparsely glandular-hairy, rarely glabrous;

corolla pale blue or pale lilac, rarely white, 2.5–5 mm diam.;

stamens 2 mm;

style 0.9–2 mm.

Capsules

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

slightly compressed in cross section, obcordiform to globular, 2–3 × 2–3 mm, apex rounded or ± emarginate, sparsely glandular-hairy.

Seeds

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

30–40, ochre, broadly ellipsoid, ± flat, convex on both sides, 0.5–0.6 × 0.4–0.5 mm, thickness and texture unknown.

2n

= 18.

= 54 (Asia).

Veronica copelandii

Veronica undulata

Phenology Flowering Jul–Aug. Flowering (Mar–)Apr–Sep.
Habitat Subalpine meadows, alpine slopes. Disturbed, wet places, ditches, or swamps.
Elevation 1900–2500 m. (6200–8200 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
AL; OR; WA; Asia [Introduced in North America]
[BONAP county map]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the flora area, Veronica undulata was introduced near ports (Mobile, Alabama, and Portland, Oregon) before 1900 via ship ballast from trade with Asia, did not spread much, and may not have persisted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 310. FNA vol. 17, p. 314.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. verna, V. wormskjoldii
Name authority Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi) Wallich: in W. Roxburgh, Fl. Ind. 1: 147. (1820)
Web links