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Copeland's speedwell

false ivy-leaf speedwell

Habit Perennials. Annuals.
Stems

ascending, unbranched, 5–15 cm, densely glandular-hairy.

decumbent to ascending, sometimes erect in young plants, 5–40(–50) cm, eglandular-hairy.

Leaves

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

blade ovate to broadly ovate, 5(–7)-lobed, central lobe longer than wide and broadest, (7–)9–15(–25) × (7–)9–17(–27) mm, base truncate, margins 5(–7)-lobed, apex obtuse to rounded, surfaces sparsely eglandular-hairy.

Racemes

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

1–10, terminal, 50–400(–500) mm, 5–20-flowered, axis eglandular-hairy;

bracts broadly ovate, 5(–7)-lobed, central lobe longer than wide and broadest, (7–)9–15(–25) mm.

Pedicels

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

patent or deflexed, (7–)10–18(–24) mm, ± longer than subtending bract, length 3–4(–6) times calyx, eglandular-hairy (densely adaxially).

Flowers

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

calyx lobes 3–4 mm, apex acute, sparsely hairy on abaxial surface or glabrous, ciliate with 25–35 hairs per side;

corolla pale violet to whitish, 4–5(–6) mm diam.;

stamens pale violet, 0.4–0.8 mm;

style 0.2–0.7 mm.

Capsules

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

± round in cross section, ovoid, 2–3 × 3.5–5 mm, apex ± emarginate, glabrate.

Seeds

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

1–4, bright reddish brown, broadly ovoid to subglobular, urn-shaped, 2.2–2.7 × 2–2.4 mm, 1–1.8 mm thick, transverse ribs inconspicuous, thinner towards margin, margins strongly involute.

2n

= 18.

= 36 (Europe).

Veronica copelandii

Veronica sublobata

Phenology Flowering Jul–Aug. Flowering Mar–May.
Habitat Subalpine meadows, alpine slopes. Moist forests, damp, shady ruderal places, thickets, gardens, montane fields.
Elevation 1900–2500 m. (6200–8200 ft.) 0–1000 m. (0–3300 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
AL; KY; MD; MO; OH; UT; VA; Europe [Introduced in North America]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica sublobata is closely related to V. hederifolia and has been frequently synonymized with or treated as a subspecies of the latter. It has rarely been reported in the flora area and is surely more widespread than indicated above. It seems to be more rare than V. hederifolia with only about a dozen verified records from seven states. However, its frequent occurrence in cultivated habitats in Europe makes it likely that it has been introduced to the flora area on multiple occasions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 310. FNA vol. 17, p. 315.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. hederifolia subsp. lucorum, V. hederifolia var. lucorum
Name authority Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi) M. A. Fischer: Oesterr. Bot. Z. 114: 201, 227, figs. 3c, 4c. (1967)
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