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Copeland's speedwell

creeping speedwell, slender speedwell, thread-stalk speedwell, véronique filiforme

Habit Perennials. Perennials.
Stems

ascending, unbranched, 5–15 cm, densely glandular-hairy.

creeping, 5–50 cm, eglandular- and glandular-hairy.

Leaves

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

blade suborbiculate, 4–13 × 4–14 mm, base cordate, margins crenate-serrate, teeth (2 or)3–5(or 6) per side, apex rounded, surfaces sparsely eglandular- and/or glandular-hairy.

Racemes

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

1–6, terminal, 50–500 mm, 10–20-flowered, axis sparsely eglandular- and/or glandular-hairy;

bracts suborbiculate, 4–13 mm.

Pedicels

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

patent or recurved, (15–)20–30 mm, length 2–5 times subtending bract, eglandular- and glandular-hairy.

Flowers

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

calyx lobes 4–7 mm, 1.5–2.5 mm wide, apex rounded, sparsely glandular-hairy to glabrate;

corolla blue-violet to pale blue and white, 8–14 mm diam.;

stamens 2.1–2.5 mm;

style 3–4 mm.

Capsules

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

compressed in cross section, subglobular, 3.5–5 × 5.5–6.5 mm, apex emarginate, sinus angle 50–90°, absent or almost smooth, glabrous or glandular-ciliate.

Seeds

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

(2–)6–14(–18), ochre, ovoid, flat to ± excavate, 1.3–1.7 × 1–1.3 mm, 0.6–0.8 mm thick, rugose to rugulose.

2n

= 18.

= 14 (Europe).

Veronica copelandii

Veronica filiformis

Phenology Flowering Jul–Aug. Flowering (Mar–)Apr–Jun(–Aug).
Habitat Subalpine meadows, alpine slopes. Moist lawns and meadows, waste fields, stream banks.
Elevation 1900–2500 m. (6200–8200 ft.) 0–1000 m. (0–3300 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; CT; DC; DE; ID; MD; MI; NY; OH; OR; PA; VT; WA; WV; BC; ON; QC; Eurasia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica filiformis is self-incompatible. In the flora area, plants seldom produce capsules; it is dispersed by stem cuttings. The species is native to subalpine meadows in the Caucasus and northern Turkey.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 310. FNA vol. 17, p. 320.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi) Smith: Trans. Linn. Soc. London 1: 195. (1791)
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