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Copeland's speedwell

Dillenius' speedwell

Habit Perennials. Annuals.
Stems

ascending, unbranched, 5–15 cm, densely glandular-hairy.

erect to ascending, (8–)10–20(–40) cm, glandular- and eglandular-hairy.

Leaves

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

blade lanceolate to ovate, 7–19(–21) × (3–)5–12 mm, 1–2 times as long as wide, base cuneate, margins (proximal) crenate-dentate or (distal) ±palmatifid, lobes 3–7+, apex of central lobe obtuse, lateral ± acute, surfaces glandular- and eglandular-hairy.

Racemes

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

1, terminal, 60–180 mm, 15–50(–120)-flowered, axis eglandular- and glandular-hairy;

proximal bracts similar to leaves, distal ones linear-lanceolate, 3–5 mm.

Pedicels

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

erect, 2–4(–5) mm, shorter than subtending bract, length 1/2–1 times calyx, eglandular- and glandular-hairy.

Flowers

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

calyx lobes 3.5–6 mm, apex acute, eglandular- and glandular-hairy;

corolla deep blue, 4–6 mm diam.;

stamens 0.8–1.5 mm;

style 0.8–1.5 mm, stigma violet.

Capsules

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

compressed in cross section, narrowly obcordiform, 3.5–4.5 × 4–6 mm, apex emarginate, angle of sinus ca. 90°, ciliate with glandular and eglandular hairs.

Seeds

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

10–28, yellowish, ellipsoid, flat, 0.9–1.6 × 0.7–1.3 mm, 0.2–0.3 mm thick, smooth.

2n

= 18.

= 16 (Eurasia).

Veronica copelandii

Veronica dillenii

Phenology Flowering Jul–Aug. Flowering (Mar–)Apr–Jul.
Habitat Subalpine meadows, alpine slopes. Open pine and oak forests, rocky, dry, sandy slopes.
Elevation 1900–2500 m. (6200–8200 ft.) 500–2200 m. (1600–7200 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
IL; IN; MI; NY; VA; WI; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica dillenii is closely related to V. verna but with larger flowers; it may have been overlooked and may be distributed more widely. Most herbarium specimens of V. dillenii blacken when dry due to the presence of aucubin, which distinguishes them from V. verna.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 310. FNA vol. 17, p. 318.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi) Crantz: Stirp. Austr. Fasc. ed. 2, 2: 352. (1769)
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