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Copeland's speedwell

field speedwell, green field speedwell, green procumbent speedwell, véronique agreste

Habit Perennials. Annuals.
Stems

ascending, unbranched, 5–15 cm, densely glandular-hairy.

creeping to decumbent, 5–25 cm, hairy.

Leaves

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

blade elliptic-ovate to oblong, (6–)8–16(–19) × (3–)4–10(–13) mm, base truncate, margins serrate, teeth 4–6(–8) per side, apex obtuse to acute, abaxial surface ± more densely hairy than adaxial.

Racemes

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

1–5, terminal, 50–250 mm, 5–15-flowered, axis eglandular-hairy, sometimes glandular-hairy;

bracts elliptic-ovate to oblong, (6–)9–16(–19) mm.

Pedicels

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

deflexed, (5–)6–10(–15) mm, ± shorter than subtending bracts, densely eglandular- and, sometimes, glandular-hairy distally.

Flowers

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

calyx lobes linear-lanceolate, (5–)6–7 mm, 2.2–2.6 mm wide, apex rounded, sparsely eglandular- or glandular-hairy;

corolla white or pale pinkish or pale blue, 4–5(–6) mm diam.;

stamens 0.5–1 mm;

style (0.6–)0.9–1.1(–1.2) mm.

Capsules

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

± compressed in cross section, ± subglobular, 3.5–4.5(–4.7) × 4.5–6(–6.3) mm, apex emarginate, sparsely glandular-hairy.

Seeds

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

6–14, yellow to ochre, globular, cymbiform, 1.3–2.1 × 1–1.6 mm, 0.6–1.1 mm thick, strongly rugose (transversely ribbed).

2n

= 18.

= 28 (Europe).

Veronica copelandii

Veronica agrestis

Phenology Flowering Jul–Aug. Flowering May–Oct.
Habitat Subalpine meadows, alpine slopes. Light, noncalcareous, moist soils, gravelly soils.
Elevation 1900–2500 m. (6200–8200 ft.) (0–)300–800 m. ((0–)1000–2600 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
DC; FL; IL; LA; MA; NY; PA; TX; VT; NF; NS; QC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica agrestis is similar to the more frequent V. persica and probably frequently overlooked and to be expected elsewhere. However, it should be noted that it is less common than V. persica or V. polita and always in more humid habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 310. FNA vol. 17, p. 320.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi) Linnaeus: Sp. Pl. 1: 13. (1753)
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