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chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell

undulate speedwell, wavy-leaf water speedwell

Habit Annuals or perennials. Annuals or perennials.
Stems

erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally.

erect or prostrate basally, 10–100 cm, glabrous at least proximally, glandular-hairy distally.

Leaves

petiole 0 mm;

blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous.

petiole 0–5 mm;

blade elliptic to ovate, sometimes ovate-oblong or linear-lanceolate, rarely lanceolate, 20–50(–100) × 5–20(–25) mm, 2.5–4 times as long as wide, base attenuate, upwards amplexicaul, margins subentire or crenate to serrate or ± undulate, apex acute to obtuse, surfaces glabrous.

Racemes

10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid;

pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous;

bracts oblong, 3–5 mm, apex obtuse.

6–25, axillary, 50–220 mm, 10–15 mm diam., 10–100-flowered, axis sparsely glandular-hairy, rarely glabrous;

pedicels 2–6 per cm;

bracts linear to lanceolate, 3–6 mm, apex acute.

Pedicels

patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy.

patent, 3–5 mm, equal to subtending bract, sparsely glandular-hairy.

Flowers

calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy;

corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.;

stamens 5 mm;

style (1.3–)1.5–2.5 mm.

calyx lobes 1.5–2(–3) mm, apex acute, sparsely glandular-hairy, rarely glabrous;

corolla pale blue or pale lilac, rarely white, 2.5–5 mm diam.;

stamens 2 mm;

style 0.9–2 mm.

Capsules

± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy.

slightly compressed in cross section, obcordiform to globular, 2–3 × 2–3 mm, apex rounded or ± emarginate, sparsely glandular-hairy.

Seeds

26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

30–40, ochre, broadly ellipsoid, ± flat, convex on both sides, 0.5–0.6 × 0.4–0.5 mm, thickness and texture unknown.

2n

= 36.

= 54 (Asia).

Veronica catenata

Veronica undulata

Phenology Flowering Jun–Oct. Flowering (Mar–)Apr–Sep.
Habitat Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels. Disturbed, wet places, ditches, or swamps.
Elevation 0–2500 m. (0–8200 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
[WildflowerSearch map]
from FNA
AL; OR; WA; Asia [Introduced in North America]
[BONAP county map]
Discussion

Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the flora area, Veronica undulata was introduced near ports (Mobile, Alabama, and Portland, Oregon) before 1900 via ship ballast from trade with Asia, did not spread much, and may not have persisted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 314. FNA vol. 17, p. 314.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. verna, V. wormskjoldii
Synonyms V. catenata var. glandulosa, V. connata subsp. glaberrima
Name authority Pennell: Rhodora 23: 37. (1921) Wallich: in W. Roxburgh, Fl. Ind. 1: 147. (1820)
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