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chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell

trilobed speedwell

Habit Annuals or perennials. Annuals.
Stems

erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally.

decumbent, (5–)10–30 cm, ± eglandular-hairy.

Leaves

petiole 0 mm;

blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous.

blade broadly ovate to suborbiculate, 3(–5)-lobed, central lobe 1/4–1/2 times length of whole leaf, (3–)5–10(–15) × (4–)6–12(–18) mm, ± fleshy, base truncate to subcordate, margins 3(–5)-lobed, apex acute, surfaces sparsely eglandular-hairy.

Racemes

10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid;

pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous;

bracts oblong, 3–5 mm, apex obtuse.

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis sparsely eglandular-hairy;

bracts broadly ovate to suborbiculate, 3(–5)-lobed, (3–)5–10(–15) mm.

Pedicels

patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy.

patent, deflexed in fruit, (2–)4–8(–10) mm, shorter than subtending bract, length 1–2 times calyx, glabrous abaxially, hairy adaxially.

Flowers

calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy;

corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.;

stamens 5 mm;

style (1.3–)1.5–2.5 mm.

calyx lobes 4–5(–6) mm, apex acute, puberulent on abaxial surface, rarely glabrous, densely ciliate with 35–60 per side;

corolla deep blue with white center, 3–5 mm diam.;

stamens dark blue, length unknown;

style 0.5–0.9(–1.1) mm.

Capsules

± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy.

± round in cross section, ovoid, 2.5–3(–3.5) × 4–5 mm, apex ± emarginate, glabrous.

Seeds

26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

(1–)4, dull yellowish brown, ellipsoid, often elongate-cuboidal, deeply urn-shaped (elaiosome inside), 2.2–2.8 × 1.5–2.2 mm, 1–1.4 mm thick, cristate to strongly rugose at margins.

2n

= 36.

= 18 (Eurasia).

Veronica catenata

Veronica triloba

Phenology Flowering Jun–Oct. Flowering (Feb–)Mar–May(–Jun).
Habitat Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels. Disturbed sites.
Elevation 0–2500 m. (0–8200 ft.) 100–200 m. (300–700 ft.)
Distribution
from FNA
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
[WildflowerSearch map]
from FNA
CA; OH; Europe (Balkans); Asia (Turkey) [Introduced in North America]
Discussion

Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica triloba is known in the flora area from three recent collections in California and Ohio; it is possibly more widespread than reported here. It is closely related to V. hederifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 314. FNA vol. 17, p. 316.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. catenata var. glandulosa, V. connata subsp. glaberrima V. hederifolia var. triloba
Name authority Pennell: Rhodora 23: 37. (1921) (Opiz) Opiz: Naturalientausch 11: 467. (1826)
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