Veronica catenata |
Veronica copelandii |
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chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell |
Copeland's speedwell |
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Habit | Annuals or perennials. | Perennials. |
Stems | erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally. |
ascending, unbranched, 5–15 cm, densely glandular-hairy. |
Leaves | petiole 0 mm; blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous. |
blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy. |
Racemes | 10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid; pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous; bracts oblong, 3–5 mm, apex obtuse. |
1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy; bracts lanceolate, 3–5 mm. |
Pedicels | patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy. |
erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy. |
Flowers | calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy; corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.; stamens 5 mm; style (1.3–)1.5–2.5 mm. |
calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy; corolla pale blue to purple, rotate, 8–10 mm diam.; stamens 4–5 mm; style 7 mm. |
Capsules | ± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy. |
compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy. |
Seeds | 26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown. |
2n | = 36. |
= 18. |
Veronica catenata |
Veronica copelandii |
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Phenology | Flowering Jun–Oct. | Flowering Jul–Aug. |
Habitat | Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels. | Subalpine meadows, alpine slopes. |
Elevation | 0–2500 m. (0–8200 ft.) | 1900–2500 m. (6200–8200 ft.) |
Distribution |
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
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CA |
Discussion | Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 314. | FNA vol. 17, p. 310. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. catenata var. glandulosa, V. connata subsp. glaberrima | |
Name authority | Pennell: Rhodora 23: 37. (1921) | Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi) |
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