Veronica catenata |
Veronica argute-serrata |
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chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell |
bilobed speedwell, sawtooth speedwell, twolobed speedwell |
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Habit | Annuals or perennials. | Annuals. |
Stems | erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally. |
erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy. |
Leaves | petiole 0 mm; blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous. |
blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate. |
Racemes | 10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid; pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous; bracts oblong, 3–5 mm, apex obtuse. |
1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy; bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire. |
Pedicels | patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy. |
erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy. |
Flowers | calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy; corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.; stamens 5 mm; style (1.3–)1.5–2.5 mm. |
calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy; corolla mostly intense blue with white center, 4–6 mm diam.; stamens 2.5 mm; style (0.7–)1–1.3(–1.7) mm. |
Capsules | ± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy. |
slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy. |
Seeds | 26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate. |
2n | = 36. |
= 18, 42 (Asia). |
Veronica catenata |
Veronica argute-serrata |
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Phenology | Flowering Jun–Oct. | Flowering Apr–Jun. |
Habitat | Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels. | Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils. |
Elevation | 0–2500 m. (0–8200 ft.) | (600–)900–3000 m. ((2000–)3000–9800 ft.) |
Distribution |
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
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CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
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Discussion | Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 314. | FNA vol. 17, p. 321. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. catenata var. glandulosa, V. connata subsp. glaberrima | V. bornmuelleri |
Name authority | Pennell: Rhodora 23: 37. (1921) | Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata) |
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