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chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell

American brooklime, American speedwell, véronique d'amérique

Habit Annuals or perennials. Perennials.
Stems

erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally.

decumbent or ascending, 5–50 cm, glabrous.

Leaves

petiole 0 mm;

blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous.

petiole 2–6(–10) mm;

blade ovate to lanceolate, widest proximal to middle, (5–)30–50(–100) × (3–)7–20(–30) mm, 2–4 times as long as wide, base truncate, rounded or almost cordate, abruptly turning into petioles, margins entire or serrate, apex obtuse-orbicular or acute, surfaces glabrous.

Racemes

10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid;

pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous;

bracts oblong, 3–5 mm, apex obtuse.

1–8, axillary, 30–100 mm, (3–)10–30-flowered, axis glabrous;

bracts lanceolate, 3–6 mm.

Pedicels

patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy.

patent, 3–10(–12) mm, equal to ± longer than subtending bract, glabrous.

Flowers

calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy;

corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.;

stamens 5 mm;

style (1.3–)1.5–2.5 mm.

calyx lobes 2–5(–6) mm, apex acute, glabrous;

corolla pink or sky blue, 4–10 mm diam.;

stamens 2–4 mm;

style (1.7–)2.5–3.5(–4) mm.

Capsules

± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy.

slightly compressed in cross section, globular, 2–4 × 3–5 mm, apex not or ± emarginate, glabrous.

Seeds

26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

20–30, brownish, ellipsoid to ovoid, indistinct-compressed or planoconvex, 0.5–0.7 × 0.3–0.6 mm, thickness varies due to compression in capsule, indistinctly wrinkled or ± rugose.

2n

= 36.

= 36.

Veronica catenata

Veronica americana

Phenology Flowering Jun–Oct. Flowering May–Aug.
Habitat Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels. Slowly flowing waters, banks, sand bars, gravel flood plains, moist soils, springs, ditches, swamps, marshes.
Elevation 0–2500 m. (0–8200 ft.) 0–3600 m. (0–11800 ft.)
Distribution
from FNA
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
[WildflowerSearch map]
from FNA
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Mexico; Asia (Japan, Russia)
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although difficult to separate, Veronica beccabunga and V. americana are maintained as species here. Evidence suggesting separate species status is different ploidy level (the tetraploid level is rare in V. beccabunga) and the occurrence of V. americana in more natural habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 314. FNA vol. 17, p. 313.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. catenata var. glandulosa, V. connata subsp. glaberrima V. beccabunga var. americana
Name authority Pennell: Rhodora 23: 37. (1921) Schweinitz ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 468. (1846)
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