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chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell

tail kittentails

Habit Annuals or perennials.
Stems

erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally.

Leaves

petiole 0 mm;

blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous.

persistent, some withering in 2d year as new leaves expand;

blade orbiculate to reniform, 25+ mm wide, not leathery, base cordate to lobate, margins ± incised-crenate to dentate, teeth apices rounded to acute or acuminate, surfaces glabrous, rarely sparsely hairy;

basal veins extending into distal 1/2 of blade, lateral veins 2–4 on each side of midvein.

Racemes

10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid;

pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous;

bracts oblong, 3–5 mm, apex obtuse.

erect, to 33 cm in fruit;

sterile bracts 3+, ovate-spatulate, largest 1+ cm;

flowers 12–100, loosely aggregated.

Pedicels

patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy.

Flowers

calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy;

corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.;

stamens 5 mm;

style (1.3–)1.5–2.5 mm.

Stamens

epipetalous.

Ovaries

ovules 10–16.

Capsules

± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy.

glabrous or glabrescent.

Seeds

26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

Sepals

4.

Petals

(3 or)4(or 5), apex entire or erose;

corolla blue, ± regular, campanulate, much longer than calyx, glabrous or puberulent, tube conspicuous.

2n

= 36.

Veronica catenata

Synthyris missurica

Phenology Flowering Jun–Oct.
Habitat Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels.
Elevation 0–2500 m. (0–8200 ft.)
Distribution
from FNA
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
[WildflowerSearch map]
from FNA
CA; ID; MT; OR; WA
[WildflowerSearch map]
Discussion

Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Racemes usually 20–100-flowered, to 33 cm at end of flowering; ovaries glabrous; capsules glabrous.
S. missurica subsp. major
1. Racemes usually 12–45-flowered, to 25 cm at end of flowering; ovaries sparsely hairy, especially along margins; capsules glabrescent.
→ 2
2. Leaf blade margins regularly toothed; racemes to 20 cm at end of flowering.
S. missurica subsp. missurica
2. Leaf blade margins regularly to irregularly toothed; racemes to 25 cm at end of flowering.
S. missurica subsp. stellata
Source FNA vol. 17, p. 314. FNA vol. 17, p. 300.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Synthyris
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
S. alpina, S. borealis, S. bullii, S. canbyi, S. cordata, S. dissecta, S. laciniata, S. lanuginosa, S. missurica, S. oblongifolia, S. pinnatifida, S. plantaginea, S. platycarpa, S. ranunculina, S. reniformis, S. ritteriana, S. rubra, S. schizantha, S. wyomingensis
Subordinate taxa
S. missurica subsp. major, S. missurica subsp. missurica, S. missurica subsp. stellata
Synonyms V. catenata var. glandulosa, V. connata subsp. glaberrima Veronica missurica
Name authority Pennell: Rhodora 23: 37. (1921) (Rafinesque) Pennell: Proc. Acad. Nat. Sci. Philadelphia 85: 89. (1933)
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