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bilobed speedwell, twolobe speedwell

undulate speedwell, wavy-leaf water speedwell

Habit Annuals. Annuals or perennials.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

erect or prostrate basally, 10–100 cm, glabrous at least proximally, glandular-hairy distally.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

petiole 0–5 mm;

blade elliptic to ovate, sometimes ovate-oblong or linear-lanceolate, rarely lanceolate, 20–50(–100) × 5–20(–25) mm, 2.5–4 times as long as wide, base attenuate, upwards amplexicaul, margins subentire or crenate to serrate or ± undulate, apex acute to obtuse, surfaces glabrous.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

6–25, axillary, 50–220 mm, 10–15 mm diam., 10–100-flowered, axis sparsely glandular-hairy, rarely glabrous;

pedicels 2–6 per cm;

bracts linear to lanceolate, 3–6 mm, apex acute.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

patent, 3–5 mm, equal to subtending bract, sparsely glandular-hairy.

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx lobes 1.5–2(–3) mm, apex acute, sparsely glandular-hairy, rarely glabrous;

corolla pale blue or pale lilac, rarely white, 2.5–5 mm diam.;

stamens 2 mm;

style 0.9–2 mm.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

slightly compressed in cross section, obcordiform to globular, 2–3 × 2–3 mm, apex rounded or ± emarginate, sparsely glandular-hairy.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

30–40, ochre, broadly ellipsoid, ± flat, convex on both sides, 0.5–0.6 × 0.4–0.5 mm, thickness and texture unknown.

2n

= 28 (Asia).

= 54 (Asia).

Veronica biloba

Veronica undulata

Phenology Flowering Apr–Jul. Flowering (Mar–)Apr–Sep.
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. Disturbed, wet places, ditches, or swamps.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; OR; WA; Asia [Introduced in North America]
[BONAP county map]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the flora area, Veronica undulata was introduced near ports (Mobile, Alabama, and Portland, Oregon) before 1900 via ship ballast from trade with Asia, did not spread much, and may not have persisted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 314.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. verna, V. wormskjoldii
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) Wallich: in W. Roxburgh, Fl. Ind. 1: 147. (1820)
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