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bilobed speedwell, twolobe speedwell

false ivy-leaf speedwell

Habit Annuals. Annuals.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

decumbent to ascending, sometimes erect in young plants, 5–40(–50) cm, eglandular-hairy.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

blade ovate to broadly ovate, 5(–7)-lobed, central lobe longer than wide and broadest, (7–)9–15(–25) × (7–)9–17(–27) mm, base truncate, margins 5(–7)-lobed, apex obtuse to rounded, surfaces sparsely eglandular-hairy.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

1–10, terminal, 50–400(–500) mm, 5–20-flowered, axis eglandular-hairy;

bracts broadly ovate, 5(–7)-lobed, central lobe longer than wide and broadest, (7–)9–15(–25) mm.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

patent or deflexed, (7–)10–18(–24) mm, ± longer than subtending bract, length 3–4(–6) times calyx, eglandular-hairy (densely adaxially).

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx lobes 3–4 mm, apex acute, sparsely hairy on abaxial surface or glabrous, ciliate with 25–35 hairs per side;

corolla pale violet to whitish, 4–5(–6) mm diam.;

stamens pale violet, 0.4–0.8 mm;

style 0.2–0.7 mm.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

± round in cross section, ovoid, 2–3 × 3.5–5 mm, apex ± emarginate, glabrate.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

1–4, bright reddish brown, broadly ovoid to subglobular, urn-shaped, 2.2–2.7 × 2–2.4 mm, 1–1.8 mm thick, transverse ribs inconspicuous, thinner towards margin, margins strongly involute.

2n

= 28 (Asia).

= 36 (Europe).

Veronica biloba

Veronica sublobata

Phenology Flowering Apr–Jul. Flowering Mar–May.
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. Moist forests, damp, shady ruderal places, thickets, gardens, montane fields.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 0–1000 m. (0–3300 ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; KY; MD; MO; OH; UT; VA; Europe [Introduced in North America]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica sublobata is closely related to V. hederifolia and has been frequently synonymized with or treated as a subspecies of the latter. It has rarely been reported in the flora area and is surely more widespread than indicated above. It seems to be more rare than V. hederifolia with only about a dozen verified records from seven states. However, its frequent occurrence in cultivated habitats in Europe makes it likely that it has been introduced to the flora area on multiple occasions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 315.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. hederifolia subsp. lucorum, V. hederifolia var. lucorum
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) M. A. Fischer: Oesterr. Bot. Z. 114: 201, 227, figs. 3c, 4c. (1967)
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