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bilobed speedwell, twolobe speedwell

Steller's speedwell

Habit Annuals. Perennials.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

erect to ascending, unbranched, (5–)10–20(–25) cm, eglandular-hairy.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

blade ovate, 15–30 × 10–20 mm, base rounded, margins dentate or serrate, apex subacute to subobtuse, surfaces eglandular-hairy.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

1, terminal, 10–60 mm, to 100 mm in fruit, 5–15-flowered, axis hairy;

bracts lanceolate, 2–5 mm.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

erect, 2–11 mm, equal to longer than subtending bract, crisp-hairy.

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx lobes 4–5 mm, eglandular-ciliate, apex acuminate, glabrous or sparsely hairy;

corolla pale blue or violet, rotate, 8–10 mm diam.;

stamens 7 mm;

style 3–6 mm.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

compressed in cross section, oblong, 5–6(–8) × 3–4.5 mm, longer than wide, apex ± emarginate, glabrous proximally, hairy distally.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

number unknown, brown, ovoid, flat, 1–1.5 × 0.7–1.2 mm, thickness and texture unknown.

2n

= 28 (Asia).

= 18 (Japan, Russia).

Veronica biloba

Veronica stelleri

Phenology Flowering Apr–Jul. Flowering Jul–Sep.
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. Dry to moderately moist slopes, meadows in alpine regions, moraines.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; Asia (Japan, Korea, Russia)
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica stelleri may be difficult to differentiate from 4b. V. wormskjoldii subsp. nutans where sympatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. stelleri var. glabrescens, V. wormskjoldii var. stelleri
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) Link: Jahrb. Gewächsk. 1(3): 40. (1820)
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