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bilobed speedwell, twolobe speedwell

thyme-leaf speedwell

Habit Annuals. Perennials.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

creeping to ascending, 5–40 cm, scattered eglandular hairs only, often also with glandular hairs, sometimes glabrate.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

blade oblong-lanceolate to ovate, 8–25 × 5–13 mm, 1.5–2.5 times as long as wide, base cuneate, margins subentire or serrulate-crenate, apex rounded to short-acuminate, surfaces glabrate.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

1, terminal, 50–100 mm, 10–30(–60)-flowered, axis eglandular- and glandular-hairy;

bracts oblong, 4–7 mm.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

erect, 2–5 mm, 4–6 mm in fruit, shorter than subtending bract in flower, eglandular- and, sometimes, glandular-hairy.

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx lobes 2–3 mm, apex acute, ciliate;

corolla white, blue, or pink, with purple or dark blue veins (except on abaxial lobe), rotate, 5–8 mm diam.;

stamens 2.5–3 mm;

style 2–4 mm.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

compressed in cross section, obcordiform, 2.5–3.5 × 4–5.5 mm, wider than long, apex acutely emarginate, glandular-ciliate, otherwise glabrate.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

50–72, brown, ellipsoid, flat, 0.6–1.1 × 0.4–1 mm, 0.2 mm thick, smooth.

2n

= 28 (Asia).

= 14.

Veronica biloba

Veronica serpyllifolia

Phenology Flowering Apr–Jul. Flowering Apr–Aug.
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. River banks, wet places, moist meadows, shady forests.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 0–3300 m. (0–10800 ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NL; NS; ON; PE; QC; SK; South America; Mexico (Baja California); Eurasia; Australia
[WildflowerSearch map]
[BONAP county map]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Alpine plants of Veronica serpyllifolia with bright blue, larger corollas, and smaller, hairier raceme axes have been treated at various ranks under the epithet humifusa. The characters are labile and show intergradations with the typical plants. Veronica serpyllifolia is widespread; it is not clear where it is native.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 312.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. humifusa, V. serpyllifolia var. decipiens, V. serpyllifolia subsp. humifusa, V. serpyllifolia var. humifusa, V. tenella
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) Linnaeus: Sp. Pl. 1: 12. (1753)
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