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bilobed speedwell, twolobe speedwell

bird's-eye speedwell, bird-eye speedwell, large field or bird's-eye speedwell, Persian speedwell, véronique de perse

Habit Annuals. Annuals.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

creeping to decumbent, 10–50(–60) cm, eglandular-hairy.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

blade suborbiculate, broadly ovate, or broadly lanceolate, (6–)9–18(–30) × (5–)8–15(–20) mm, base truncate, margins serrate, apex acute, surfaces sparsely eglandular-hairy.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

1–6, terminal, 100–500(–600) mm, 5–30-flowered, axis eglandular-hairy;

bracts suborbiculate or broadly ovate or broadly lanceolate, (6–)9–18(–25) mm.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

spreading, deflexed in fruit, (12–)15–27(–38) mm, length 1–2(–3) times subtending bract, densely eglandular-hairy.

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx lobes (4.5–)5.5–8(–9.5) mm, (1.7–)2.4–3.6(–4.2) mm wide, apex acuminate, eglandular-hairy;

corolla intense bright blue, 8–14 mm diam.;

stamens 1.2 mm;

style (1.5–)2–2.8(–3.2) mm.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

compressed in cross section, broadly obcordiform, 4–6 × (5–)6–8.5(–9.5) mm, apex acute, sinus angle (80–)90–120(–150)°, reticulate with prominent veins, ± sparsely to densely eglandular- and/or glandular-hairy or glabrate.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

(10–)12–18(–20), pale brownish yellow, ellipsoid to globular, cymbiform, (1.3–)1.4–2.3(–2.5) × (0.8–)0.9–1.6(–1.9) mm, 0.5–1 mm thick, cristate-rugose.

2n

= 28 (Asia).

= 28.

Veronica biloba

Veronica persica

Phenology Flowering Apr–Jul. Flowering (Jan–)Apr–Jul(–Dec).
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. Roadsides, lawns, fields, waste places.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 0–500(–2000) m. (0–1600(–6600) ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; sw Asia [Introduced in North America; introduced also in Mexico (Michoacán, Veracruz), Central America, South America, Eurasia, e Asia, Africa, Atlantic Islands, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The names Veronica buxbaumii Tenore and V. tournefortii C. C. Gmelin (not V. ×tournefortii Villars) have been used for V. persica.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 319.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. persica var. aschersoniana, V. persica var. corrensiana, V. tournefortii subsp. aschersoniana, V. tournefortii subsp. corrensiana
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) Poiret: in J. Lamarck et al., Encycl. 8: 542. (1808)
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