Veronica biloba |
Veronica officinalis |
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bilobed speedwell, twolobe speedwell |
common gypsyweed, common speedwell, heath speedwell, herbal speedwell, Paul's betony, véronique officinale |
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Habit | Annuals. | Perennials. |
Stems | erect, (1–)2–10(–18) cm, eglandular-hairy. |
creeping, decumbent to ascending distally, 10–40(–50) cm, densely eglandular-hairy, rarely glabrate or glabrous. |
Leaves | blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate. |
blade ovate, ovate-oblong, or obovate, (8–)20–35(–50) × (3–)10–20(–30) mm, 1.5–2 times as long as wide, base attenuate to cuneate or obtuse, margins dentate, denticulate, or serrulate, apex obtuse to rounded, surfaces sparsely to densely eglandular-hairy, rarely glabrate. |
Racemes | 1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy; bracts lanceolate, 6–9 mm, margins entire, apex acute. |
1–4(–8), axillary, 40–60 mm, 10–35(–40)-flowered, axis densely to sparsely eglandular-hairy, sometimes also glandular-hairy, rarely glabrate; bracts linear-lanceolate or narrowly ovate, (1.5–)3–4(–5) mm. |
Pedicels | straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous. |
erect, (0.5–)1–2(–3) mm, shorter than subtending bract, densely eglandular-hairy, rarely glandular-hairy or glabrate. |
Flowers | calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate; corolla pale to clear blue, 2–4 mm diam.; stamens 1–2 mm; style 0.4–0.8(–2) mm. |
calyx 4(or 5)-lobed, lobes 2–3.5(–4) mm, apex obtuse, glandular- and eglandular-hairy; corolla pale blue-lilac to pale pink, rarely white with pink veins, 6–8 mm diam.; stamens 2.8–3.4 mm; style 2.5–3(–4) mm. |
Capsules | slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous. |
compressed in cross section, triangulate-obdeltoid to obcordiform, 3–5 × 4–5 mm, apex rounded or truncate, rarely emarginate, glandular-hairy, rarely glabrous. |
Seeds | (2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth. |
9–24, brown to yellow, ellipsoid, ovoid, or obovoid, flat, 0.8–1.6 × (0.3–)0.6–1.2 mm, 0.2–0.3 mm thick, smooth. |
2n | = 28 (Asia). |
= 18, 36 (Europe). |
Veronica biloba |
Veronica officinalis |
|
Phenology | Flowering Apr–Jul. | Flowering (Apr–)Jun–Jul(–Aug). |
Habitat | Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. | Forests, meadows, pastures, scrub, ruderal places. |
Elevation | (400–)1300–3000 m. ((1300–)4300–9800 ft.) | 0–1600 m. (0–5200 ft.) |
Distribution |
MT; UT; BC; Asia [Introduced in North America]
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CA; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Eurasia [Introduced in North America; introduced in s South America (Chile)]
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Discussion | Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters. Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica ×tournefortii Villars [V. officinalis var. tournefortii (Villars) Dumortier] refers to the hybrid between V. officinalis and V. allionii Villars in Europe and is not applicable to any North American plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 321. | FNA vol. 17, p. 311. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Name authority | Linnaeus: Mant. Pl. 2: 172. (1771) | Linnaeus: Sp. Pl. 1: 11. (1753) |
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