Veronica biloba |
Veronica longifolia |
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bilobed speedwell, twolobe speedwell |
long-leaf speedwell, véronique à longues feuilles |
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Habit | Annuals. | Perennials. |
Stems | erect, (1–)2–10(–18) cm, eglandular-hairy. |
erect, usually with 10–20 nodes, (50–)70–120(–150) cm, proximal 1/2 glabrous, distal 1/2 eglandular-hairy with some glandular hairs intermixed. |
Leaves | blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate. |
blade narrowly ovate-lanceolate, almost linear, or narrowly ovate, (60–)80–120(–150) × 10–30(–35) mm, base cuneate, truncate, or cordate, margins serrate to biserrate, apex acute, surfaces sparsely or densely eglandular-hairy, rarely glabrous. |
Racemes | 1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy; bracts lanceolate, 6–9 mm, margins entire, apex acute. |
1–7(–9), terminal and occasionally axillary, 100–200(–300) mm, 150–300-flowered, axis eglandular- and glandular-hairy; bracts linear to subulate, 2–6 mm. |
Pedicels | straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous. |
erect, (1–)1.5–3(–5) mm, shorter than subtending bract, densely eglandular-hairy, sometimes also glandular-hairy. |
Flowers | calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate; corolla pale to clear blue, 2–4 mm diam.; stamens 1–2 mm; style 0.4–0.8(–2) mm. |
calyx lobes 2–2.5(–4) mm, apex acute, eglandular-hairy, sometimes glandular-hairy; corolla blue, campanulate, longer than wide, 5–7 mm diam.; stamens 5–6 mm; style (5–)6–8(–10) mm. |
Capsules | slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous. |
± compressed in cross section, ovoid, 2.5–3 × 2.5–3 mm, apex emarginate, glabrous. |
Seeds | (2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth. |
2–40, light brown, ellipsoid, flat, 0.7–1.2 × 0.5–0.8 mm, 0.2–0.3 mm thick, smooth. |
2n | = 28 (Asia). |
= 34, 68 (Eurasia). |
Veronica biloba |
Veronica longifolia |
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Phenology | Flowering Apr–Jul. | Flowering Jun–Aug(–Sep). |
Habitat | Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. | Moist (swampy) habitats, roadsides. |
Elevation | (400–)1300–3000 m. ((1300–)4300–9800 ft.) | 0–800 m. (0–2600 ft.) |
Distribution |
MT; UT; BC; Asia [Introduced in North America]
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AK; CT; FL; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; ND; NH; NJ; NY; OH; PA; RI; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Eurasia [Introduced in North America; introduced also in South America, Pacific Islands (New Zealand)]
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Discussion | Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters. Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Reports of Veronica bachofenii Heuffel, V. grandis Fischer ex Sprengel (= V. daurica Steven), or V. spuria Linnaeus, seem always to refer to garden varieties of V. longifolia. Garden varieties often differ from typical plants in having glandular hairs and broader leaves, which may indicate past hybridization with other species. Subgenus Pseudolysimachium includes Veronica longifolia and V. spicata, which are naturalized in the flora area, but also other species and hybrids of these, which are commonly grown ornamentally and may escape from cultivation. All of these plants should easily key out with V. longifolia and V. spicata but differ in specifics of the species descriptions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 321. | FNA vol. 17, p. 308. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. maritima | |
Name authority | Linnaeus: Mant. Pl. 2: 172. (1771) | Linnaeus: Sp. Pl. 1: 10. (1753) |
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