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bilobed speedwell, twolobe speedwell

rock speedwell, woodystem speedwell

Habit Annuals. Perennials.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

ascending to erect, densely branched from woody base, (5–)10–15(–30) cm, eglandular-hairy, sometimes glandular-hairy.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

blade obovate to ovate or spatulate, sometimes suborbiculate proximally, (7–)8–20(–25) × (2–)3–6(–7) mm, mostly shorter to equal to internodes, base cuneate, margins entire or ± crenate or serrate, apex acute, surfaces glabrous or glabrate.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

1–10(–20), terminal, rarely with 1–4 axillary, 20–40 mm, (1–)4–10(–18)-flowered, axis glabrate to sparsely eglandular-hairy;

bracts linear or linear-lanceolate or long-ovate, (1–)3.5–9(–15) mm.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

erect, (2–)5–7(–15) mm, equal to ± longer than subtending bracts, eglandular-hairy.

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx 4(or 5)-lobed, lobes (2.5–)4.5–6(–8) mm, apex obtuse, eglandular-hairy, sometimes also glandular-hairy;

corolla intense blue, sometimes with reddish or white center, rarely white, rotate, (6–)10–14(–15) mm diam.;

stamens 5–7 mm;

style (3–)5–7(–8.5) mm.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

± compressed in cross section, ovoid, (5–)6–8(–9) × (3–)4–4.5(–5.5) mm, longer than wide, apex attenuate to apiculate and acute, rarely rounded, not emarginate, eglandular-hairy, sometimes mixed with larger glandular hairs.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

35–62, orange, ellipsoid, flat, 0.9–1.5 × (0.4–)0.9–1.2 mm, 0.2–0.3 mm thick, smooth.

2n

= 28 (Asia).

= 16.

Veronica biloba

Veronica fruticans

Phenology Flowering Apr–Jul. Flowering (May–)Jun–Aug.
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. Fissures, rocky places, scree.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 0–1000 m. (0–3300 ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from USDA
Greenland
[BONAP county map]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 318.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) Jacquin: Enum. Stirp. Vindob. 2: 200. (1762)
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