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bilobed speedwell, twolobe speedwell

Dillenius' speedwell

Habit Annuals. Annuals.
Stems

erect, (1–)2–10(–18) cm, eglandular-hairy.

erect to ascending, (8–)10–20(–40) cm, glandular- and eglandular-hairy.

Leaves

blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate.

blade lanceolate to ovate, 7–19(–21) × (3–)5–12 mm, 1–2 times as long as wide, base cuneate, margins (proximal) crenate-dentate or (distal) ±palmatifid, lobes 3–7+, apex of central lobe obtuse, lateral ± acute, surfaces glandular- and eglandular-hairy.

Racemes

1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy;

bracts lanceolate, 6–9 mm, margins entire, apex acute.

1, terminal, 60–180 mm, 15–50(–120)-flowered, axis eglandular- and glandular-hairy;

proximal bracts similar to leaves, distal ones linear-lanceolate, 3–5 mm.

Pedicels

straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous.

erect, 2–4(–5) mm, shorter than subtending bract, length 1/2–1 times calyx, eglandular- and glandular-hairy.

Flowers

calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate;

corolla pale to clear blue, 2–4 mm diam.;

stamens 1–2 mm;

style 0.4–0.8(–2) mm.

calyx lobes 3.5–6 mm, apex acute, eglandular- and glandular-hairy;

corolla deep blue, 4–6 mm diam.;

stamens 0.8–1.5 mm;

style 0.8–1.5 mm, stigma violet.

Capsules

slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous.

compressed in cross section, narrowly obcordiform, 3.5–4.5 × 4–6 mm, apex emarginate, angle of sinus ca. 90°, ciliate with glandular and eglandular hairs.

Seeds

(2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth.

10–28, yellowish, ellipsoid, flat, 0.9–1.6 × 0.7–1.3 mm, 0.2–0.3 mm thick, smooth.

2n

= 28 (Asia).

= 16 (Eurasia).

Veronica biloba

Veronica dillenii

Phenology Flowering Apr–Jul. Flowering (Mar–)Apr–Jul.
Habitat Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. Open pine and oak forests, rocky, dry, sandy slopes.
Elevation (400–)1300–3000 m. ((1300–)4300–9800 ft.) 500–2200 m. (1600–7200 ft.)
Distribution
from FNA
MT; UT; BC; Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
IL; IN; MI; NY; VA; WI; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters.

Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica dillenii is closely related to V. verna but with larger flowers; it may have been overlooked and may be distributed more widely. Most herbarium specimens of V. dillenii blacken when dry due to the presence of aucubin, which distinguishes them from V. verna.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 318.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Linnaeus: Mant. Pl. 2: 172. (1771) Crantz: Stirp. Austr. Fasc. ed. 2, 2: 352. (1769)
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