Veronica biloba |
Veronica catenata |
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bilobed speedwell, twolobe speedwell |
chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell |
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Habit | Annuals. | Annuals or perennials. |
Stems | erect, (1–)2–10(–18) cm, eglandular-hairy. |
erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally. |
Leaves | blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate. |
petiole 0 mm; blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous. |
Racemes | 1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy; bracts lanceolate, 6–9 mm, margins entire, apex acute. |
10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid; pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous; bracts oblong, 3–5 mm, apex obtuse. |
Pedicels | straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous. |
patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy. |
Flowers | calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate; corolla pale to clear blue, 2–4 mm diam.; stamens 1–2 mm; style 0.4–0.8(–2) mm. |
calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy; corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.; stamens 5 mm; style (1.3–)1.5–2.5 mm. |
Capsules | slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous. |
± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy. |
Seeds | (2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth. |
26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
2n | = 28 (Asia). |
= 36. |
Veronica biloba |
Veronica catenata |
|
Phenology | Flowering Apr–Jul. | Flowering Jun–Oct. |
Habitat | Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. | Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels. |
Elevation | (400–)1300–3000 m. ((1300–)4300–9800 ft.) | 0–2500 m. (0–8200 ft.) |
Distribution |
MT; UT; BC; Asia [Introduced in North America]
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CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
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Discussion | Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters. Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 321. | FNA vol. 17, p. 314. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. catenata var. glandulosa, V. connata subsp. glaberrima | |
Name authority | Linnaeus: Mant. Pl. 2: 172. (1771) | Pennell: Rhodora 23: 37. (1921) |
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