Veronica biloba |
Veronica arvensis |
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bilobed speedwell, twolobe speedwell |
corn speedwell, speedwell, véronique des champs, wall speedwell |
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Habit | Annuals. | Annuals. |
Stems | erect, (1–)2–10(–18) cm, eglandular-hairy. |
erect to ascending, 1–30(–40) cm, glandular- or eglandular-hairy. |
Leaves | blade ovate to lanceolate-ovate, (3–)4–12(–20) × (1–)2–6(–12) mm, base cuneate, margins remotely serrate, rarely dentate, apex acute, surfaces glabrate. |
in 3–6 pairs per stem; blade oblong to broadly ovate, (2–)5–14(–35) × (2–)3–10(–18) mm, 2.5 times as long as wide, base truncate, margins crenate-serrate, apex obtuse to rounded, surfaces sparsely eglandular-hairy, rarely also glandular-hairy. |
Racemes | 1–8, terminal, 20–80 mm, 5–15(–30)-flowered, axis eglandular- and glandular-hairy; bracts lanceolate, 6–9 mm, margins entire, apex acute. |
1(–3), terminal, 20–150 mm, (5–)15–40(–60)-flowered, axis densely eglandular- and glandular-hairy; bracts linear-oblong, (3.5–)4–6(–10) mm. |
Pedicels | straight or scarcely deflexed, erect-spreading to horizontally spreading in fruit, (2–)3–8(–11) mm, shorter than subtending bract, eglandular-hairy, sometimes glandular-hairy, rarely glabrous. |
erect, 0–4 mm, shorter than subtending bract, densely eglandular- and glandular-hairy. |
Flowers | calyx lobes (3–)4–7(–9) mm, margins entire, apex acuminate-acute, pairs connate 0.5–1.5 mm, puberulent or glabrate; corolla pale to clear blue, 2–4 mm diam.; stamens 1–2 mm; style 0.4–0.8(–2) mm. |
calyx lobes (2–)3.5–5(–6) mm, 0.8–2 mm wide, apex acute, glandular-hairy; corolla sky blue to intense blue, 2–4 mm diam.; stamens 0.3–0.5 mm; style (0.2–)0.4–0.6(–1) mm. |
Capsules | slightly compressed in cross section, obcordiform, (2.4–)2.8–4(–4.5) × (3–)3.5–5(–6) mm, apex emarginate, eglandular- and/or glandular-hairy, rarely glabrous. |
compressed in cross section, obcordiform, 2–4 × 2.5–5 mm, apex markedly emarginate, glandular-ciliate, otherwise glabrous. |
Seeds | (2–)4–6(–10), bright yellow, oblong, cymbiform, (1.4–)1.7–2(–2.2) × (0.7–)1–1.3 mm, 0.7 mm thick, smooth. |
10–30, yellowish, ovoid, flat, 0.7–1.7 × 0.4–1 mm, 0.2–0.4 mm thick, rugose. |
2n | = 28 (Asia). |
= 16. |
Veronica biloba |
Veronica arvensis |
|
Phenology | Flowering Apr–Jul. | Flowering (Feb–)Mar–Jun(–Oct). |
Habitat | Bare soils, open forests, scree, mountain and alpine meadows, humid places near streams, waste fields, ruins, calcareous and non-calcareous soils. | Disturbed sites, lawns, fields, open forests, scrub, grasslands, rocky sites, coasts. |
Elevation | (400–)1300–3000 m. ((1300–)4300–9800 ft.) | 0–2900 m. (0–9500 ft.) |
Distribution |
MT; UT; BC; Asia [Introduced in North America]
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AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; YT; Atlantic Islands; Pacific Islands; Greenland; Africa; Eurasia; Australia [Introduced in North America; introduced also in Mexico, Central America]
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Discussion | Most reports of Veronica biloba pertain to V. argute-serrata. There are gatherings from potentially mixed populations (for example, Mt. Baldy, British Columbia, Lomer 97-118, UBC). Studies on phenotypic plasticity in these two species are necessary to ascertain the diagnostic characters. Veronica intercedens Bornmüller, native to western Asia, was recently reported from Idaho. It is most similar to V. biloba, distinguished by a large calyx with broadly ovate lobes (3–5 mm versus 2–3.5 mm), the pairs joined by more than one half. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 17, p. 321. | FNA vol. 17, p. 317. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Name authority | Linnaeus: Mant. Pl. 2: 172. (1771) | Linnaeus: Sp. Pl. 1: 13. (1753) |
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