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American brooklime, American speedwell, brooklime, European speedwell, véronique beccabunga

Cusick's speedwell

Habit Perennials.
Stems

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 18, 72.

Veronica beccabunga

Veronica cusickii

Phenology Flowering Jun–Aug.
Habitat Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
CA; CT; IL; MA; MD; ME; MI; MO; NJ; NV; NY; OH; PA; VA; WI; WV; BC; ON; QC; Eurasia; Africa [Introduced in North America; introduced also in s South America]
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from FNA
CA; ID; MT; OR; WA; BC
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Discussion

Subspecies 3 (1 in the flora).

Subspecies abscondita M. A. Fischer is found from eastern Turkey to western Iran, while the native range of subsp. muscosa (Korshinsky) Elenevsky extends from eastern Iran to southern China.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 312. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Subordinate taxa
V. beccabunga subsp. beccabunga
Synonyms V. allenii
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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