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American brooklime, American speedwell, brooklime, European speedwell, véronique beccabunga

Copeland's speedwell

Habit Perennials.
Stems

ascending, unbranched, 5–15 cm, densely glandular-hairy.

Leaves

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

Racemes

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

Pedicels

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

Flowers

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

Capsules

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

Seeds

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

2n

= 18.

Veronica beccabunga

Veronica copelandii

Phenology Flowering Jul–Aug.
Habitat Subalpine meadows, alpine slopes.
Elevation 1900–2500 m. (6200–8200 ft.)
Distribution
from FNA
CA; CT; IL; MA; MD; ME; MI; MO; NJ; NV; NY; OH; PA; VA; WI; WV; BC; ON; QC; Eurasia; Africa [Introduced in North America; introduced also in s South America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 3 (1 in the flora).

Subspecies abscondita M. A. Fischer is found from eastern Turkey to western Iran, while the native range of subsp. muscosa (Korshinsky) Elenevsky extends from eastern Iran to southern China.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 312. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Subordinate taxa
V. beccabunga subsp. beccabunga
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi)
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