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corn speedwell, speedwell, véronique des champs, wall speedwell

Copeland's speedwell

Habit Annuals. Perennials.
Stems

erect to ascending, 1–30(–40) cm, glandular- or eglandular-hairy.

ascending, unbranched, 5–15 cm, densely glandular-hairy.

Leaves

in 3–6 pairs per stem;

blade oblong to broadly ovate, (2–)5–14(–35) × (2–)3–10(–18) mm, 2.5 times as long as wide, base truncate, margins crenate-serrate, apex obtuse to rounded, surfaces sparsely eglandular-hairy, rarely also glandular-hairy.

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

Racemes

1(–3), terminal, 20–150 mm, (5–)15–40(–60)-flowered, axis densely eglandular- and glandular-hairy;

bracts linear-oblong, (3.5–)4–6(–10) mm.

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

Pedicels

erect, 0–4 mm, shorter than subtending bract, densely eglandular- and glandular-hairy.

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

Flowers

calyx lobes (2–)3.5–5(–6) mm, 0.8–2 mm wide, apex acute, glandular-hairy;

corolla sky blue to intense blue, 2–4 mm diam.;

stamens 0.3–0.5 mm;

style (0.2–)0.4–0.6(–1) mm.

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

Capsules

compressed in cross section, obcordiform, 2–4 × 2.5–5 mm, apex markedly emarginate, glandular-ciliate, otherwise glabrous.

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

Seeds

10–30, yellowish, ovoid, flat, 0.7–1.7 × 0.4–1 mm, 0.2–0.4 mm thick, rugose.

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

2n

= 16.

= 18.

Veronica arvensis

Veronica copelandii

Phenology Flowering (Feb–)Mar–Jun(–Oct). Flowering Jul–Aug.
Habitat Disturbed sites, lawns, fields, open forests, scrub, grasslands, rocky sites, coasts. Subalpine meadows, alpine slopes.
Elevation 0–2900 m. (0–9500 ft.) 1900–2500 m. (6200–8200 ft.)
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; YT; Atlantic Islands; Pacific Islands; Greenland; Africa; Eurasia; Australia [Introduced in North America; introduced also in Mexico, Central America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 13. (1753) Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi)
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