Veronica arvensis |
Veronica anagallis-aquatica |
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corn speedwell, speedwell, véronique des champs, wall speedwell |
blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
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Habit | Annuals. | Annuals or perennials. |
Stems | erect to ascending, 1–30(–40) cm, glandular- or eglandular-hairy. |
erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
Leaves | in 3–6 pairs per stem; blade oblong to broadly ovate, (2–)5–14(–35) × (2–)3–10(–18) mm, 2.5 times as long as wide, base truncate, margins crenate-serrate, apex obtuse to rounded, surfaces sparsely eglandular-hairy, rarely also glandular-hairy. |
petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
Racemes | 1(–3), terminal, 20–150 mm, (5–)15–40(–60)-flowered, axis densely eglandular- and glandular-hairy; bracts linear-oblong, (3.5–)4–6(–10) mm. |
6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
Pedicels | erect, 0–4 mm, shorter than subtending bract, densely eglandular- and glandular-hairy. |
suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
Flowers | calyx lobes (2–)3.5–5(–6) mm, 0.8–2 mm wide, apex acute, glandular-hairy; corolla sky blue to intense blue, 2–4 mm diam.; stamens 0.3–0.5 mm; style (0.2–)0.4–0.6(–1) mm. |
calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
Capsules | compressed in cross section, obcordiform, 2–4 × 2.5–5 mm, apex markedly emarginate, glandular-ciliate, otherwise glabrous. |
slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
Seeds | 10–30, yellowish, ovoid, flat, 0.7–1.7 × 0.4–1 mm, 0.2–0.4 mm thick, rugose. |
40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
2n | = 16. |
= 36. |
Veronica arvensis |
Veronica anagallis-aquatica |
|
Phenology | Flowering (Feb–)Mar–Jun(–Oct). | Flowering Mar–Sep(–Nov). |
Habitat | Disturbed sites, lawns, fields, open forests, scrub, grasslands, rocky sites, coasts. | Stream margins, ditches, banks, springs, swamps, wet meadows. |
Elevation | 0–2900 m. (0–9500 ft.) | 0–4000 m. (0–13100 ft.) |
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; YT; Atlantic Islands; Pacific Islands; Greenland; Africa; Eurasia; Australia [Introduced in North America; introduced also in Mexico, Central America]
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AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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Discussion | Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 17, p. 317. | FNA vol. 17, p. 313. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera | |
Name authority | Linnaeus: Sp. Pl. 1: 13. (1753) | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) |
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